RETTORI, ROBERTO

SENOWBARI-DARYAN, BABA

ZÜHLKE, RAINER

English

ROBERTO RETTORI

BABA SENOWBARI-DARYAN

RAINER ZÜHLKE

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<EM>FLATSCHKOFELIA ANISICA</EM> GEN. ET SP. N. (FORAMINIFERIDA) FROM THE MIDDLE TRIASSIC (ANISIAN) OF NORTHERN DOLOMITES, ITALY

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Rivista Italiana di Paleontologia e Stratigrafia volume 102 numero 3 tavola 1 pagrne 413-476 Dicembre 1996FLATSCHKOFELIA ANISICA GEN. ET SP. NOV. (FORAMINIFERIDA)FROM THE MIDDLE TRIASSIC (ANISIAN) OF NORTHERN DOLOMITES, ITALYROBERTO RETTORI*. BABA SENOWBARI.DARYAN'S* & RAINER ZUHLKE***Key-utords: Paleontologl Foraminifers, Triassic, Anisian, Alps,Nonhern Dolomites, Italy.Riassunto. Nel presente lavoro viene descritto un nuovo gener€di foraminifero sessile a parete agglutinante: Flatscbkofelia (specie-tipo:Flatschkofelia anisica gen et sp. nov.). Questo foraminifero è stato rin-venuto nelle brecce di talus recifale di età Triassico medio (Anisico)de1la Formazione del Serla superiore (Dolomiti di Valdora, Italia).Flatscbhofelia, gen. nov., è riferito alla famiglia Placopsilinidae Rhum-b1er.Abstract. The new genus of sessile and agglutinated foramini-fer, Flaxschhofelia (type-spectes: Flatschhofelia anisica gen. et sp. nov.)from the Middle Triassic (Anisian) reef talus of the Olang Dolomites(lJpper Serla Formation, Dolomiti di Valdora, Italy) is introduced.Flaxcbkofelia, gen. nov., is referred to the family PlacopsilinidaeRhumbler.lntroduction.After the end-Permian mass extinction, reefs andreefal carbonate buildups did not exist during the LatePermian to Early Triassic. First occurrence of reefs hasbeen reported from the Early-Middle Anisian (Middleand Late Pelsonian) of the Dolomites, Italy and fromSouthern China (see Senowbari-Daryan er al., 1993). Inthe Northern part of the western Tethys additional Ani-sian reefs are known from the Carpathians and from theKarawanken Mountains. From the southern paft of thewestern Tethys no Anisian reefs are known.Most of Late Anisian reef carbonates in the OlangDolomites (Southern Alps, Italy) are represented by al-lochthonous talus reef blocks of one or several cubicme-ter size. Autochthonous reef buildups are extremelyrare. Both, the allochthonous reef blocks and autochtho-nous buildups occur within the Middle to Late Pelso-nian Recoaro Formation. The fossil content as well asthe microfacies types of these reef carbonates are descri-bed by Bechstàdt 6c Brandner (1.970), Fois & Gaetani(198a) and by Senowbari-Daryan et al. (1993) in detail.Olangocoelia ozr Bechstadt & Brandner, a problematicfossil (sponge or alga?), is the most abundant organismin the Anisian reef carbonates, followed by different ty-pes of sphinctozoid and inozoid sponges, bryozoans, so-lenoporacean algae, corals, encrusting epibionts (poro-stromate algae, cyanophycean crusts, microproblematica,worms and foraminifers) (see for detaii Senowbari-Da-ryan et a1., 1,993). Foraminifers are relatively rare andmostly represented by Meandrospira dinarica Ko-chansky-Devidè & Pantic, Pilammina densa Pantic, En-doteba sp., Endotebanella sp., Endouiada sp., Endotria-della sp., Glomospira sp., Glomospirella sp. Duostomini-dae etc. Sessile foraminifers are also not abundant andrepresented by the questionable gena,s Bullopora? andbyan agglutinated type, described here as Flatschhofelia ani-slca gen. et sp. nov.Systematic descriptionForaminiferida Eichwald, 1830Suborder Text u I a ri i n a Delage & Hérouard, 1896Superfamily L i t u o I a c ea de BlainviTle, 1827Family Placopsilinidae Rhumbler, 1913Subfamily Placopsilininae Rhumbler, 191.3Genus Flatschkofelia gen. nov.Type-species: Flatschkofelia anisica gen. et sp. nov.Derivation of name. After the locality Flatschkofel, OlangDolomites, Italy (Fig. 1), where the holotype was found.Diagnosis. Test simple, agglutinated and solidstructured, attached on the early coiled stage, free anduncoiled on the final stage. Chambers are biserially ar-'r Dipanimento di Scienze della Terra, Università degli Studi di Perugia, Piazza Università 4, 06100 Perugìa, Italy.'r'r Institut fùr Palaontologie, Universitàt Erlangen-Nùrnberg, LoewenichstraBe 28, D-91054 Erlangen, Germany.ooo Geologisch-Palàontologisches Institut, Universitàt Heidelberg, Im NeuenheimerFeld234,D-69120 Heidelberg, Germany./'"\:--' )--r-- )-^ì-v;fir*.--'; /ANAB@enMamolada o.-L--30*FurkeLPmPiz da Per6  DreifrngerspitzeLocalitY,É Maurerkopf^^Flatschkofel O12|lrnR. Rettori, B. Senoubari.Dartan €r R. ZùblkeFig. 1 - Location map of the Dolomites and Flatschkofel, whereFlatscbkofelia anisica gen. et sp. nov. was found.ranged in the initial stage, bi- and uniserial in later stage.Aperture simple and terminal located in the first biserialstage, moving to the margin in the second uniserial stage.Composition of the genus. At the moment mono-typic, only represented by the type-species.Stratigraphic and geographic distribution. MiddleTriassic (Anisian), Northern Dolomites, Italy.Remarks. Flatscbkofelia gen. nov. is herein inclu-ded in the family Placopsilinidae Rhumbler, 1913 deil-ned by I-oeblich & Tappan (1987, p. 80) as follows:"Test attached, early stage coiled, arcuate, or biserial, la-ter uncoiled; wall agglutinated, solid, aperture terminal,single or multiple. L. Jurassic to Flolocene"Tho f"-il-' i..l'161s5 two subfamilies: Placopsilini-nae Rhumblen I9I3 and Adhaerentiinae Loeblich EcTappan, L986. Flatscbkofelia gen. nov., is referred to Pla-copsilininae because having a coiled early stage even ifall the genera belonging to the subfamily lacks a biserialPaft.The biserial arrangement of the chambers is amorphoiogical feature of Adhaerentiinae, but the uniquegenus (Adhaerentia Plummer, 1938, type-species Adhae-rentia midaayensis Plummer, 1938) included in thesubfamily lacks an early coiled stage.The presence of a coiled early stage, a biserial se-cond part and a final uniserial part, which is commonboth in Placopsilininae and Adhaerentiinae, rogerherwith aperturai features, could justify the introduction ofa new subfamily for Flatschkofelia, gen. nov. Flowever,the material from the Northern Dolomites is notenough to introduce a new subfamily.At the momenr, Flatscbkofelia is referred to Pla-copsiiinidae also because the Lower Paleocene genusAdhaerentia shows, in the final srage, aperrure multiplewith two to many separate openings and for strati-graphic reasons too.According to Loeblich & Tappan (1987) the strati-graphic range of Placopsilinidae spans from I-ower Juras-sic to Holocene, but we can affirm the range of the fa-mily starts from Triassic, nor only on rhe basis of theexistence of the new genus Flatschkofelia, bur also becau-se of previous record of the genus Placopsilina in Trias-sic sediments (Trifonova 1967, 1992; Brónnimann & Za-nínettt. 1972\.Flatschkofelia anisica gen. et sp. nov.PL 1., Íig. 1-121993 Sessile agglutinated {oraminifer (gen. et sp. indet.) - Senowbari-Daryan, Ztihlke, Bechstid & Flùgel, p. 230, pl. 64, fig. 2,4,5.Derivation of name. Because rhe occurrence of the soecies in Ani-sirn limestones.Holotype. Longitudinal section illustrated in Pl. 1, fig. 1 (thin sec-tion: 89lV4,/18).Paratypes. All specirnens illustrated in Pl. 1, fig. 2-12 from differentthin sections (see explanation of plate 1).Type locality. Flatschkofel, Olang Dolomites, Italy (see Fig. 1).Type level. Upper Serla Formation, Middle Triassic (Anisian, Pelso-nran).Material. Several specimens in different planes from thin sections:89/Y7/49, 89/Vr/85, 89/V4/t8 (holotype), 89/Vr, 86/2,89/y7/32, deposited at the Institute fùr Pa1àontologie, Universitvof Erlangen-Nùrnberg.Diagnosis. See diagnosis of the genus.Italy; Middle Triassic (Anisian,8, Ia) 8e/Y4/1.8r 7) 89/Y1t e)PLATE 1Fig. 1-12 - Flatscbhofelia anisica gen. et sp. nov. from Flatschkofel N, Upper Serla Formation, Olang Dolomites,Pelsonian).Fig. 1: Holotype, thin section 89/V4/1,8. Fìg. 2-72: Paratypes, thin sections: 2) 89/V7/32i 3, 4, 5, 6,86/Vr/85r 1r, 12) 89/V1/8s.415Pl. 1 Flatschkofelia anisica gen. et sp. no'u..,,ts-O O.SmmR. Rettori, B. Senoubari-Daryan & R. ZilhlkeDescription. The test is attached, at least for theinitial part, then, probably, may grow free of the at-tachment. It is irregulariy coiled in the reduced earlystage, later uncoiled and rectilinear, slightly contortedabove aii in the final part. The coiled early stage is fol-lowed by a short stage with 2-3 series of subangularchambers biserially arranged. The final part is uniserialand the chambers (5-7 ín the more developed specimens)are irregular in shape, increasing more rapidly inbreadth than height.The aperture is simple, terminally located in thebiserial stage, moving to the margin in the uniserialpart, at the end of a short neck in any chamber offsettoward the opposite margin in respect with the previouschamber. The chamber wall is solid, agglutinated andsimple in structure.Dimensions of the test (in mm):Lensnt: u.)-1.+uMaximum diameter: 0.4Height of the chmbers in the uniserial stage: 0.06-0.08Breath of the chambers in the uniserial stage: 0.16-0.25Thickness of the *all: 0.05-0.06Stratigraphic and geographic distribution. MiddleTriassic (Anisian, Pelsonian), Northern Dolomites, Italy.Association. Flaxcbkofelia anisica, gen. et sp. nov.,was found within upper ramp and lagoonal facies of theprograding Upper Serla Formation. The underlaying Re-coaro Formation and the Upper Serla Formation containallochthonous reef facies (reef talus biocks, reworked bio-clastic respectively). For details on Anisian reef orga-nisms of Olang Dolomites see Senowbary-Daryan et aI.(1993). The species is associated with rare foraminifersmainly represented by Endoteba sp., Endotebanella sp.,Endotriada sp. and Endotriadella sp. in the reef facies;Meandrospira dinarica Kochansky-Devidè & Pantic andPilammina densa Pantíc in the lagoonal facies.Acknouled.gments.The investigations wereForschungsgemeinschaft (Pro;euPriority Program "Global andSedimentation".supported by the DeutscheBe 641/17, Anis-RifQ in theRegionel Controls on BiogenicREFERENCESBechstàdt T. & Brandner R. (1970) - Das Anis zwischen St.Vigil und dem Hóhensteintal (Pragser- und Olanger Do-lomiten, Sùdtirol). Festband Geol. Inst., 300-Jahr-Feier,Univ. Innsbruck, pp. 9-103.Brónnimann P. & Zaninetti L. (1,972) - Foraminifera from thebasal Upper Muschelkalk at Hyères, western Basse-Pro-vence, Southern France. Rio. It. Paleont. Strat., v.78, n.l, pp. 43-44, Milano.Fois E. 8r Gaetani M. (1984) - The recovery of reef-buildingcommunities and the role of cnidarians in carbonate se-quences of the Middle Triassic (Anisian) in the ItalianDolomites. Paleontograpbica Amer., v. 54, pp. l9L2A0,Ithaca/New York.Loebiich A.R. E Tappan E. (1987) - Foraminiferal Genera andtheir classification. Van Nostrand Reinhold ComoanvInc., Vol. oÎ 970 pp., New York.Senowbari-Daryan B., Zùhlke R., Bechstàd T. 6c Flùgei E.(1,993) - Anisian (Middle Triassic) Buildups of theNorthern Dolomites (Italy): the recovery of reef corn-munities after the Permian,/Triassic Crisis. Facies. v. 28,pp. 181-256, Erlangen.Trifonova E. (1964 - Some new Triassic foraminifera in Bul-garla. Ann. Unio. Sofia, Geol., Geogr., Fac. 60 (1 Geol.)pp. 5-6, Sofia.Trifonova E. (1992) - Taxonomy of bulgarian Triassic Forami-nifera. I. Families Psammosphaeridae to Nodosinelli-dae. Geol. Balc., v. 22, n. 7, pp. 3-50, Sofia.Receioed June 20, 1996; accepted October 3, 1996

FLATSCHKOFELIA ANISICA GEN. ET SP. N. (FORAMINIFERIDA) FROM THE MIDDLE TRIASSIC (ANISIAN) OF NORTHERN DOLOMITES, ITALY

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FLATSCHKOFELIA ANISICA GEN. ET SP. N. (FORAMINIFERIDA) FROM THE MIDDLE TRIASSIC (ANISIAN) OF NORTHERN DOLOMITES, ITALY

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