journal articleThe following list of the ants of Utah was prepared from specimens and data now in the Biology Department of the University of Utah. This list is by no means complete, yet it is representative of practically all parts of the state. A few specimens in the collection date back to 1902, and all of these specimens and numerous other specimens in the collection were identified by Professor W. M. Wheeler. Since 1937 the authors have made collections in all parts of the state in an effort to obtain material for this paper. In addition, Mr. Grundmann has made intensive collections and a study of the ants of Salt Lake County as thesis material for a Master's Degree

Grundmann, A. W.

Rees, Don M.

English

The University of Utah: J. Willard Marriott Digital Library

BULLETIN OF THE U N IVERSITY OF U TA HVolume 31 November 5, 1940 No. 5A  Preliminary List of the Ants of UtahB YDON M. REES and A . W . GRU N D M AN N  University of UtahBIOLOGICAL SERIES, Vol. VI, No. 2PUBLISHED BY TH E  UNIVERSITY OF U TA H  SA L T LAKE CITYT H E  U N I V E R S I T Y  P R E S S  U N I V E R S I T Y  O F  U T A H  S A L T  L A K E  C I T YA  P r e l i m i n a r y  L is t  o f  t h e  A n t s  o f  U t a h(Hymenoptera; Formicidae)ByD o n  M. R e e s  and A. W . G r u n d m a n n  University of UtahThe following list of the ants of Utah was prepared from  speci­mens and data now  in the B iology Department of the University of Utah. This list is by no means complete, yet it is representative of practically all parts of the state. A  few specimens in the collection date back to 1902, and all o f these specimens and numerous other specimens in the collection were identified by Professor W . M. Wheeler. Since 1937 the authors have made collections in all parts of the state in an effort to obtain material for this paper. In addi­tion, Mr. Grundmann has made intensive collections and a study of the ants of Salt Lake County as thesis material for a Master’s Degree.The known ants of Utah to date com prise 90 named species, subspecies and varieties. The collection localities range from  the W arm  Temperate to the Sub-Arctic zone and occupy diversified habitats from  dry barren sand dune areas to the moist shady Sub- Alpine meadows.The authors are deeply indebted to Dr. A. C. Cole, Jr., of the University of Tennessee, for his kindness in identifying material and in verifying identifications. Records of form icidae from  Utah, as published by Professor W . M. W heeler and others, have been freely used in preparing this preliminary list.In addition to the species recorded here, Eciton coecum  Latreille, Eciton (Acamatus) melsheimeiri Haldeman, and Eciton (Acamatus) harrisi Haldeman are reported by W heeler as occurring in Utah; but this is a mistake due to assuming that Fort Gates, from  where the species are recorded by Haldeman in Stansbury’s report, is in Utah, whereas it is in Texas.A LIST OF THE ANTS OF UTAH Subfamily MyrmicinaeGenus MONOMORIUM MayrMonomorium minimum BuckleyUtah: Blanding, April 28, 1928 (A. M. W o o d b u ry ); American Fork, April 11, 1931 (R ees-M offett); Salt Lake City, April 4, 1932 (J. A. R ow e); Salt Lake County, 1937-39 (Grundmann and R ees); Swasey Springs, Millard County, May 16, 1938 (R ees ).Genus SOLENOPSIS FabricusSolenopsis molesta var. validiuscula EmeryUtah: Salt Lake City, April and May, 1931 (J. A. R o w e ); Salt Lake County, 1937-38, numerous specimens (G rundm ann); Leeds, April 4, 1939 (J. V asquez); Salt Lake City, May 2, 1939 (R ees); La Sal Creek, La Sal Mts., June 9, 1939 (R ees).Genus PHEIDOLE W estw oodPheidole californica MayrUtah: Salt Lake City, 1902 (R. V. Cham berlin); Salt Lake City, April 11, 1939 (G rundm ann).Pheidole pilifera subsp. artemisia ColeUtah: Provo, May 13, 1932 (A. C. C o le ).1Genus CREMASTOGASTER LundCremastogaster lineolata near cerasi FitchUtah: Salt Lake County, 1937-39, numerous specimens (Grund­m ann); Swasey Springs, Millard Co., May 16, 1938 (R ees).Cremastogaster lineolata subsp. coarctata var. mormonum EmeryU tah : Salt Lake City and East Millcreek Canyon, Salt Lake County, 1902 (R. V. Cham berlin).10Genus APHAENOGASTER MayrAphaenogaster subteranea subsp. occidentalis EmeryUtah: Little W illow  Creek Canyon, Salt Lake Co., July 22, 1902 (R. V. C ham berlin); Salt Lake City, September 11, 1928 (W . J. G ertsch ); Salt Lake County, 1937-39, numerous specimens (Grundmann and R ees).Aphaenogaster uinta W heelerUtah: East Millcreek Canyon, Salt Lake County, 1902, type specimen (R. V. Cham berlin); Salt Lake City and vicinity, 1937-39 (Grundm ann).Genus POGONOMYRMEX MayrPogonomyrmex barbatus var. molefaciens Buckley Utah: St. George (V . M. Tanner).0Pogonomyrmex barbatus var. fuscatus EmeryUtah: Goulding Mountain, July 11, 1937 (R. Shaw ).Pogonomyrmex barbatus var. marfensis W heeler Utah: Green River, May 5, 1933 (J. A. R ow e).4  A PRELIMINARY LIST OF THE ANTS OF UTAHPOGONOMYRMEX AND MYRMICA 5P o g o n o m y r m e x  barbatus subsp. rugosus EmeryUtah: Hurricane, April 7, 1938 (Grundm ann); St. George, August 26, 1939 (R ees ).pogonomyrmex occidentalis CressonU tah : Salt Lake County, July 22, 1902 (R. V. Cham berlin); Lehi (W . A. H ooker); Sandy, Kaysville (E. H. Klam bach)8; Brigham City (Grace O lsen)0; Tooele, Grantsville, Snowville, Ogden, Kanab, Zion National Park (A. C. C o le )6; Grantsville (R. C. Shannon)fa; Beaver, May 7, 1932 (J. A. R ow e); Salt Lake and Summit Counties, 1937-39 (Grundmann and R ees); Millard County, May 8, 1938 (R ees ); Leeds, April 18, 1939 (J. V asquez); La Sal, La Sal Junction, W ales, Brush Creek, St. George, 1939 (R e e s ).Pogonomyrmex occidentalis var. utahensis CreightonUtah: Zion National Park, July 18, 1932 (W . S. C reighton)0; Kanab, June 14, St. George, June 16 and August 26, 1939 (R ees ).Pogonomyrmex californicus BuckleyUtah: St. George (V. M. Tanner)0; Hurricane, June 15, 1939 (R ees ).Genus MYRMICA LatreilleMyrmica mutica EmeryUtah: Salt Lake City, 1902 (R. V. Cham berlin); Salt Lake City, W oods Cross, Springville, Salem, 1937-39 (Grundm ann).Myrmica rubra subsp. brevinodis Emery U tah: Salt Lake City (C. E m ery).9Myrmica rubra subsp. brevinodis var. sulcinodoides Emery U tah: Salt Lake City (C. E m ery).0Myrmica scabrinodis subsp. lobocornis var. fracticornis EmeryUtah: Salt Lake County, 1902 (R. V. Cham berlin); Salt Lake County, 1937-39, numerous specimens (G rundm ann); Summit County, Soapstone Canyon, Uinta Mts., Novem ber 9, 1938 (Grundm ann); W ales, Sanpete County, July 18, 1939 (R ees); H enry’s Fork Basin, Summit County, August 12, 1939 (R ees).Myrmica scabrinodis subsp. brevinodis EmeryUtah: Big C ottonw ood Canyon, Salt Lake County, October 11,1938 (Grundm ann).Myrmica scabrinodis subsp. sulcinodoides NylanderUtah: River Heights, Cache County, July 16, 1935 (C. F. S m ith ).5Myrmica sabuleti Meinert subsp. americana W eber Utah: Bryce Canyon (Neal W eb er).76 A PRELIMINARY LIST OF THE ANTS OF UTAHGenus LEPTOTHORAX MayrLeptothorax acervorum M ayr subsp. canadensis ProvancherUtah: Salt Lake County, 1902 (R. V. Cham berlin); 1937-1939, numerous specimens (Grundm ann).Leptothorax acervorum Mayr subsp. canadensis var. yankee Emery Utah: 1903, Ac. Nat. Sci. Phila.KLeptothorax nevadensis W heelerUtah: Blacksmith Fork Canyon, June 9, 1935 (Sm ith -R ow e)/'Leptothorax nitens EmeryUtah: Am erican Fork Canyon, Utah County, type locality (T. H. Pergande)."Genus SYMM YRM ICA W heelerSymmyrmica chamberlini W heelerU tah: Salt Lake City, August, 1902, type specimens (R. V. Cham berlin).Subfamily DolichoderinaeGenus LIOMETOPUM MayrLiometopum apiculatum subsp. luctuosum W heelerUtah: Gunnison Butte, Green River, May 7, 1932 (J. A. R o w e ); M yton, April 24, 1938 (H. G o ld ) ; Moab, June 7, and Hatch W ash, near La Sal, June 10, 1939 (R ees).Genus DORYM YRM EX MayrDorymyrmex pyramicus RogerUtah: St. George, April 7, 1938, Salt Lake County, 1937-39 (G rundm ann); Kanab, June 14, Hurricane, June 15, and St. George, June 16, 1939 (R ees).Genus TAPINOM A FoersterTapinoma sessile SayUtah: South Fork of Raft River, Box Elder County, September 7, 1932 (R ees ); Salt Lake County, 1937-38 (G rundm ann); Leeds, May 4, 1939 (J. Vasquez).Genus IRIDOMYRMEX Mayr[ridomyrmex analis AndreUtah: Salt Lake County, May 29, 1938, and September 6, 1938 (Grundm ann).IRIDOMYRMEX a n d  LASIUS 7Iridomyrmex pruinosus RogerUtah: John ’s Canyon, San Juan County, May 2, 1935 (B. E. R e e s ); Hurricane, May 7, 1938, and Salt Lake County, 1937-38 (G rundm ann).Iridomyrmex pruinosus analis AndreUtah: Pelican Point, Utah County, and Salt Lake County, 1938 (G rundm ann).Subfamily Campanotinae Genus LASIUS FabricusLasius niger var. neoniger EmeryU tah: American Fork Canyon, July 4, 1938 (K. W h ite lock ); Salt Lake County, 1938-39, numerous specimens (Grundm ann).Lasius niger var. americanus EmeryUtah: Salt Lake County, 1938 (Grundm ann); Millard County, May 18, 1938 (R e e s ).Lasius niger subsp. alienus Foerster var. americanus EmeryUtah: Salt Lake County, July 22, 1902 (R. V. Cham berlin); Salt Lake County, 1938 (Grundm ann).Lasius niger var. sitkaensis PergandeUtah: Utah County, 1932 (J. A. R o w e ); Salt Lake County, 1938­39, numerous specimens (G rundm ann); La Sal, June 9, 1939 (R ees).Lasius flavus var. nearcticus W heelerUtah: Bluff, May 11, 1933 (J. A. R ow e); Butterfield Canyon, Salt Lake County, September 3, 1938 (G rundm ann); La Sal M oun­tains, June 9, 1939 (R ees ).Lasius umbratus subumbratus ViereckUtah: Salt Lake County, July 22, 1902 (R. V. Cham berlin); May2, 1939 (G rundm ann).Lasius umbratus mixtus var. aphidicola W alshU tah : Little W illow  Creek Canyon, Salt Lake County, July 22, 1902 (R. V. Cham berlin).Lasius (Acanthomyops) interjectus Mayr Utah: Cedar City (R. V. Cham berlin).Lasius (Acanthomyops) claviger RogerUtah: Lake Blanche, W asatch Mountains, Salt Lake County, August 22, 1938 (G rundm ann).Lasius (Acanthomyops) latipes W alshUtah: Salt Lake County, 1902, and M onticello, April 15, 1928 (R. V. Chamberlin).Genus FORMICA LinneFormica sanguinea subnuda EmeryUtah: Palisade Park, Ashley National Forest, August 9, 1939 (R e e s ); H enry’s Fork Basin, August 10, 1939 (R ees ); Soapstone Canyon, Uinta Mountains, Summit County, August 25, 1939 (Grundm ann).Formica sanguinea var. puberula EmeryU tah: Stockton (T. Spaulding)12; La Sal Mountains (W . S. C reighton)2; W ales, Sanpete County, July 27, 1939 (R ees).Formica manni -U tah: Leeds, May 4, 1939 (J. Vasquez).Formica near manniUtah: W asatch Mountains, Salt Lake County, 1937-38 (Grund­m ann); Leeds, May 4, 1939 (J. Vasquez).Formica perpilosa W heelerUtah: Hurricane, April 7, 1938, Stansbury Island, Great Salt Lake, July 19, 1938, Salt Lake City, April 11, 1939 (Grund­m ann); Hurricane, June 15, 1939 (R ees).Formica rufa aggerans W heelerUtah: Lehi (W . A. H ook er)11; Swasey Springs, Millard County, May 16, 1938, Green Lake, Daggett County, August 8, 1939, and' W ales, Sanpete County, July 16, 1939 (R ees).Formica rufa subsp. obscuripes ForelUtah: Salt Lake County, 1937-39, numerous specimens (Grund­mann) .Formica integroides EmeryUtah: Swasey Springs, Millard County, May 15, 1938 (R ees).Formica truncicola subsp. integroides EmeryUtah: Logan Canyon, June 9, 1935 (T. O. Thatcher).5Formica truncicola subsp. integroides var. coloradoensis WheelerUtah: Daniel’s Canyon, August 16, 1935 (K now lton) °; Swasey Springs, Millard County, May 16, 1938 (R ees ).Formica truncicola subsp. integroides var. haemorrhoidalis Emery U tah: Bryce Canyon, 1931 (J. A. R ow e). -Formica truncicola subsp. obscuriventris MayrUtah: Salt Lake County, 1938-39, numerous specimens (Grund­mann) .Formica truncicola subsp. obscuriventris var. aggerans W heeler Utah: Prom entory Point, Great Salt Lake (A. W hetm ore) .128 A PRELIMINARY LIST OF THE ANTS OF UTAH ,FORMICA 9Formica foreliana W heelerUtah: Uinta Mountains, Summit County, October 8, 1938 (G rundm ann).Formica ciliata MayrUtah: Blanding, April 17, 1928 (A. M. W oodbu ry ).Formica comata W heelerUtah: Millcreek Canyon, Salt Lake County, October 31, 1937 (G rundm ann).Formica crinoventris W heelerUtah: Boulten, September 12, 1935 (G. F. ICnowlton).3Formica oreas W heelerUtah: Blacksmith Fork, September 7, 1932 (S m ith -R ow e)5; Boxelder County, September 7, 1932 (R ees); Duck Creek, Cedar Mountains, August 6, 1935 (A . M. W o o d b u ry ).Formica oreas var. comptula W heelerUtah: St. George, April 25, 1935, and Cornish, June 16, 1935 (G. F. K n ow lton )0; W olf Creek, Summit County, August 7, 1939 (R ees ).Formica fusca LinneUtah: Ferron, 1928 (J. A. R o w e ); Uinta Mountains, Summit County, October 8, 1938 (G rundm ann); La Sal Creek, La Sal Mountains, June 9, and Kanab Canyon, June 14, 1939 (R ees).Formica microgyna rasilis W heelerUtah: Salt Lake County, 1902 (R. V. Cham berlin)11; La Sal Mountains (W . S. C reighton).11Formica fusca var. argentea W heelerUtah: Salt Lake County, 1902 (R. V. Cham berlin); Monticello, April 17, 1928 (R. V. Cham berlin); Boxelder County, April, 1932, American Fork, April 25, 1934 (R ees ); Salt Lake County, 1937-38 (G rundm ann); Sheep Creek, Ashley National Forest, August 8, 1939 (R ees).Formica fusca var. subaenescens EmeryUtah: Salt Lake County, 1902 (R. V. Cham berlin), 1931 (J. A. R ow e), April 16, 1931 (R ees), 1937-39, numerous specimens (G rundm ann); Henry’s Fork Basin, Summit County, and Green Lake, Daggett County, August, 1939 (R ees).Formica fusca var. gelida W heelerU tah: Salt Lake County, 1938-39 (Grundm ann).Formica fusca var. neorufibarbis EmeryUtah: Salt Lake County, April 10, 1931 (R e e s ); Salt Lake County, 1937-39, numerous specimens, W oods Cross, Novem ber 7, 1937, W ellsville, September 5, 1938, and Soapstone Canyon, Uinta Mountains, October 8, 1938 (G rundm ann); Ephraim, July12, 1939 (R e e s ); Parley’s Canyon, Salt Lake County, August13, 1939 (Rees).10 A PRELIMINARY LIST OF THE ANTS OF UTAHFormica rufibarbis var. gnava Buckley U tah: Lehi, (W . A. H ook er).11Formica cinerea MayrUtah: W asatch, September 1, 1937 (Knowlton-H arm ston) ,5Formica cinerea altipetens W heelerUtah: W arner Ranger Station, La Sal Mountains (W . S. Creigh­to n )2; H enry’s Fork Basin, Summit County, August 13, 1939 (R e e s ).Formica cinerea var. neocinerea W heelerUtah: W ellsville, September 5, 1938 (G rundm ann).Formica subpolita MayrUtah: Hurricane, April 7, 1938, and Salt Lake County, November7, 1938 (G rundm ann); Swasey Springs, Millard County, May 19, 1939 (R ees).Formica subpolita var. camponoticeps W heelerU tah: Moab, 1928 (R. V. Cham berlin); Salt Lake County, 1938­39 (G rundm ann); Swasey Springs, Millard County, May 18,1939 (R e e s ).Formica wheeleri CreightonUtah: W arner Ranger Station, La Sal Mountains (W . S. Creigh­ton ) , type locality.1Formica neogagates EmeryUtah: Prom entory Point (A . W h etm ore )12; Gunnison Butte, Green River, May 7, 1932 (J. A. R o w e ); Salt Lake County, numerous specimens, 1938-39 (Grundm ann).Formica neogagates subsp. lasioides var. vetula (W heeler).Utah: Gunnison Butte, Green River, May 7, 1932 (J. A. R ow e); La Sal Mountains, W arner Ranger Station (W . S. C reighton)2; Salt Lake County, October 29, 1938 (G rundm ann); Ferron Reservoir, Emery County, July 11, 1939 (R ees).Formica pallidefulva nitidiventris EmeryUtah: Parley’s Canyon, Salt Lake County, May 11, 1938 (Grund­m ann) .Formica (Neoformica) moki W heelerUtah: M ilford (J. C. B rad ley )12; Blanding, April 17, 1928 (A. M. W ood b u ry ); Bluff, April 17, 1928 (R. V. Chamberlin).Genus POLYERGUS LatreillePolyergus rufescens Latr. subsp. breviceps EmeryUtah: Richfield, 1933 (J. A. R ow e); Chester, Sanpete County, July 14, 1939 (R e e s ).MYRMECOCYSTUS AND CAMPONOTUS 11Genus MYRMECOCYSTUS W esm aelMyrmecocystus melliger subsp. mendax W heelerUtah: W ellsville, September 5, 1938 (Grundm ann).Myrmecocystus mexicanus var. horti-deorum McCookUtah: Bluff (A. M. W oodb u ry ); between Bluff and Blanding, April 17, 1928 (R. V. Cham berlin); Gunnison Butte, Green River, May 7, 1932 (J. A. R ow e).Genus CAMPONOTUS MayrCamponotus laevigatus SmithUtah: Beaver Canyon (C. Shaeffer).12Camponotus maculatus var. vicinus MayrUtah: Moab, 1928 (R. V. Cham berlin); American Fork, April 11, 1931 (R ees-M offett); Ferron, June 6, 1932 (J. A. R o w e ); Swasey Springs, Millard County, May 15, 1938 (R ees ); Salt Lake County, 1938-39, numerous specimens (G rundm ann); La Sal Creek, La Sal Mts., June 9, 1939 (R ees ).Camponotus maculatus subsp. sansabeanus BuckleyUtah: Parowan, April 6, 1938, and Salt Lake County, September9, 1938, and April 8, 1939 (G rundm ann).Camponotus maculatus subsp. sansabeanus var. torrefactus W heeler Utah: East Millcreek Canyon, July 22, 1902 (R. V. Chamberlin), co-type; Oris, 1928 (Cham berlin); Salt Lake City, 1931 (J. A. R o w e ); Salt Lake County, 1938-39, numerous specimens (Grund­mann) .Camponotus herculeanus var. whymperi ForelUtah: Salt Lake County, 1938-39, numerous specimens (Grund­mann) ; Henry’s Fork, Summit County, August 11, 1939 (Rees).Camponotus herculeanus var. modoc W heelerUtah: Salt Lake County, 1938-39, numerous specimens (Grund­mann) ; Ferron Reservoir, Emery County, July 11, and Palisade Forest Park, Daggett County, August 8, 1939 (Rees).Camponotus herculeanus subsp. ligniperda var. noveboracensis Fitch Utah: Salt Lake County, 1937-39, numerous specimens (Grund­mann) .Camponotus caryae var. decipiens EmeryUtah: East Millcreek Canyon, July 22, 1902 (R. V. Cham berlin).BIBLIOGRAPHY1. Co l e , A. C. Descriptions of tw o new ants of the genus Pheidole. Ent. Soc. Amer., Ann. 26:616-618. 1933.2. C r e i g h t o n , W . S. Descriptions of three new North American ants with certain ecological observations on previously described form s. Psyche 61:185-200. 1934.3. C r e i g h t o n , W . S. Two new species of Formica from  western United States. Amer. Mus. Nov. No. 773, Jan. 28, 1935.4. H a l d e m a n ,  S. S. Insects in appendix C to Howard Stansbury’s “ Exploration and Survey of the Valley of the Great Salt Lake of U tah,”  pp. 367-368. 1852.5. K n o w l t o n , G. F. Hymenoptera of Utah. Utah State Agri. Col. mimeographed Bui. 1939.6. O l s e n , O . W i l f r e d . Notes on the North American harvesting ants of the Genus Pogonom yrm ex Mayr. Bui. Mus. Comp. Zool. 7 7 (8 ) :493-514. 1934.7. W e b e r , N e a l . Descriptions of new North American species and subspecies of Myrmica Latreille. Lloydia 2 :144-152. 1939.8. W h e e l e r , W i l l i a m  M o r t o n . A revision of the North American ants o f the genus Leptothorax Mayr. Acad. Nat. Sci., Phila. Proc. 55:215-260. 1903.9.  . Notes on a new guest ant, Leptothorax glacialis,and the varieties of Myrmica brevinodis Emery. Bui. W isconsin Nat. Hist. Soc. 5 (2 ) :70-83. 1907.10.  . The ants of Texas, New M exico and Arizona. Bui.Am er. Mus. Nat. Hist. 24 (21):399-485 . 1908.11.  . A  revision of the ants of the genus Formica (Linne)Mayr. Bui. Mus. Comp. Zool. 53 (10) :379-565. 1913.12.  . Mountain ants of western North America. Am er.Acad. Arts & Sci. 52:457-569. 1917.

A preliminary list of the ants of Utah

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A preliminary list of the ants of Utah

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