The terrestrial biosphere is an integral component of the Earth Observing System (EOS) science objectives concerning climate change, hydrologic cycle change, and changes in terrestrial productivity. The fluxes o f CO2 and other greenhouse gases from the land surface influence the global circulation models directly, and changes in land cover change the land surface biophysical properties o f energy and mass exchange. Hydrologic cycle perturbations result from terrestrially-induced climate changes, and more directly from changes in land cover acting on surface hydrologic balances. Finally, both climate and hydrology jointly control biospheric productivity, the source o f food, fuel, and fiber for humankind. The role of the land system in each of these three topics is somewhat different, so this chapter is organized into the subtopics of Land-Climate, Land-Hydrology, and Land-Vegetation interactions (Figures 5.1, 5.2, and 5.3)