Ruckes, Herbert, 1895-

American Museum of Natural History Scientific Publications

)>btwiican%MlsdumPUBLISHED BY THE AMERICAN MUSEUM OF NATURAL HISTORYCENTRAL PARK WEST AT 79TH STREET, NEW YORK 24, N.Y.NUMBER 1702 JANUARY 14, 1955Three New Species and a Subspecies of theGenus Mozena (Heteroptera, Coreidae)BY HERBERT RUCKES'While studying several rather large collections of coreids, the authorhas come to the conclusion that a number of new species of the genusMozena may be erected at this time. In all probability additional ones willbe found as still other collections are examined and more extensive col-lecting is carried on, especially in tropical and subtropical areas.In the course of this undertaking the author has become impressed bythe fact that one North American species widely identified as M. lanceo-lata (Herrich-Schaeffer) does not at all conform to the original descrip-tion and the colored figure that Herrich-Schaeffer includes in his"Atlas." Herrich-Schaeffer definitely states that in M. lanceolata theover-all color is dark brown, the basal segment of the antennae is black(schwartz), that the humeri are bluntly angled (stumpfen Ecken), thatthe scutellum is dark with a very pale border (Rand des Schildschens amhellsten), that the connexivum is strikingly alternated (schwartz undgelb), and that the raised lines on each side of the abdominal segments areoblique (schrdgstehenden). The species that so often has heretofore beendetermined as being lanceolata is a much paler form, with sharp-angledhumeri, a pale scutellum, a vaguely and incompletely alternated connexi-vum, and with more or less straight rather than oblique laevigate lineson each abdominal segment. This species is apparently restricted to thesouthwestern part of the United States and northern Mexico. Herein Iam describing it as Mozena arizonensis.1 Research Associate, Department of Insects and Spiders, the American Museumof Natural History, and Professor Emeritus, the City College of New York.AMERICAN MUSEUM NOVITATESMozena arizonensis, new speciesOver-all color variable but tending to be olivaceous tan with a lightcastancous suffusion, lighter than in allied species. Basal three segmentsof the antennae concolorous pale orange, orange-brown, or dull yellow,terminal one dusky; antennal segmental ratios: male: 142/140/109/109;female: 110/108/91/91; i.e., segments I and II subequal, III and IVsubequal, each of the latter about one-sixth shorter than the former.Head above, pale and usually concolorous, the eyes reddish fuscous,ocelli sanguineous; a pair of small, ivory-colored tubercles behind eacheye.Pronotum irregularly punctate, producing a somewhat rugose surfacewith no evident discal tuberculations such as are frequently found inallied species. Anterolateral margins before the humeri straight, some-times very obscurely sinuate, and provided with a few inconspicuous,rather evenly spaced tubercles; humeri lunately flaring, each terminatingin an acute, almost acuminate, slightly retrose tip; posterior margin ofeach humerus serratulate.Scutellum slightly shorter than wide across the base (101 by 109),transversely rugose, and usually concolorous dull yellow; if not concolor-ous, then only obscurely darkened centrally.Hemelytra somewhat darker than adjacent parts, the discal portionwith a reddish brown tinge; the costal and apical margins dirty ivory-yellow, with a concentration of the same color apically; numerous vari-able pale stellate points scattered over the corium; hemelytral membranedark castaneous to fuscous; hind wings almost hyaline, with only a faintsmoky tinge.Connexivum narrowly exposed, alternated dark and light, but notconspicuously so; the posterior half of the segment darker, the coloringmore intensified and distinct on the posterior three or four segments, theanterior ones only vaguely alternated, if at all.Under surface of the head, sometimes the first antennal segment, theprothorax and mesothorax, the anterior and middle tibiae, and all tarsitending to be pale olivaceous, sometimes with a slight reddish tinge. Undersurface of metathorax and abdomen suffused with red, brown, or cinna-mon over a paler dirty yellow or tan background, median portion ofthe abdomen usually the palest; the raised laevigate lines of each side ofthe abdominal venter, parallel to the margins of the segment, not oblique,and collectively forming one continuous line, not an interrupted seriesof ridges on each side, as in allied species.Inner surface of the posterior femora in varying shades of castaneous,the outer surface always paler; usually three longitudinal dorsal rows2 NO. 1702RUCKES: GENUS MOZENAof small tubercles present although there is a tendency for the ectal rowto become obsolescent, particularly in the female. At least the proximalhalf (in the males much more) of the inner surface of the hind tibiae darkfuscous to piceous, distal portion paler; the outer surface always palerand sometimes with an olivaceous tinge.The upper surface of the abdomen, as seen after the wings are spread,muddy orange in color, with variable blotching and suffusion of duskybrown which is usually concentrated posteriorly.Male genital segment similar to that in allied species, but contrastinglypaler than the adjacent abdominal plates.Described from 80 specimens.HOLOTYPE: Male, 21 mm. long; 8.75 mm. across humeri; 7.0 mm.wide through abdomen. Patagonia, Santa Cruz County, Arizona, July23, 1937. Deposited in the American Museum of Natural History.ALLOTYPE: Female, 19 mm. long; 8.0 mm. wide across humeri; 7.4mm. wide through abdomen. Tucson, Pima County, Arizona, June 6,1952. Deposited in the American Museum of Natural History.PARATYPES: Arizona: Maricopa County, Phoenix, April 16, 15 males,six females; Yavapai County, no date, two males, two females; GilaCounty: Globe, no date, one male, one female; Ash Creek, August 15,1920, two males; Roosevelt, August 15, 1920, one female; Sierra AnchaMountains, no date, one male; Pima County, Sabino Canyon, June 6,1952, three males, five females; Tucson, June 6, 1952, three males, twofemales; San Vicente, June 10, 1952, one male, five females; Redington,September 1, 1950, one female; Pinal County, Florence, July 27, 1952,two males; Oracle, July 11, 1940, one female; Chandler, July 16, 1940,three males; Cochise County, Wilcox, September 8, 1950. one male;Coconino County, Flagstaff (Oak Creek Canyon), July 27, 1950, onemale.Sonora: Pitiquito, July 4, 1952, four males, five females; Pesqueira,July 20, 1952, one male; Naco, August 15, 1949, one male; Magdalena,July 23, 1952, two males, one female; Minas Nuevas, August 7, 1952,one male, one female; Navajoa, August 2, 1952, one male. Chihuahua:Ojo Laguna, June 30, 1947, one male. Baja California: El Mayor, June15, 1952, one male.All paratypes are deposited in the American Museum of NaturalHistory.As indicated above, there has been some confusion in the proper deter-mination and naming of this species. The flaring, sharp-pointed humerisuggest relationship to M. lunata (Burmeister), but the pale over-allcolor in contrast with the very dark fuscous shades exhibited by lunata1955 3AMERICAN MUSEUM NOVITATESdefinitely set it off from that species. The muddy admixture of orange-brown and fuscous seen on the abdominal dorsum seems to show linkageto M. obtusa Uhler, where a similar coloring occurs. Very definitely thespecies just described is not the true M. lineolata (Herrich-Schaeffer).The general body proportions are somewhat like those of lunata.Mozena tomentosa, new speciesA relatively robust species with a rather widely flaring abdomen; some-what in general structure like M. lineolata (Herrich-Schaeffer). Abovefuscous, with paler markings as specified below; below dull reddishbrown, not the bright cinnamon-red found commonly in allied species;an over-all appressed tomentum present, frequently interspersed withsome fine erect setae.Head squarish, as usual, disc dark fuscous to piceous bordered witha lighter shade of brown; eyes dark brown, ocelli dull red; no postoculartubercles evident. Antennae reddish brown, with the two middle seg-ments slightly lighter, the over-all length of the antennae proportionatelyshorter than found in other species of this genus; segmental ratios:110/95/80/90.Pronotal surface somewhat undulant, with a recognizable broad grooveand shallow in the median line, posteriorly; anteriorly there are numerousirregularly placed, small, blunt tubercles on the disc, especially in theregion of the collar; humeri exceptionally large, lunately flaring, slightlyupturned and terminating in an acute point; anterolateral margin of thepronotum rather short, less than half the median length of the pronotumand provided with pronounced conical tubercles, each terminating in anerect seta; several shorter setigerous tubercles on the anterior marginof each humerus; posterior humeral margin irregularly and shallowlyerose but not serratulate.Scutellum as wide across the base as long (120 by 120); puncturescoarser and more widely spaced than on pronotum, giving a vague rugoseappearance to the surface; disc fuscous, with a narrow pale yellow border.Hemelytra more or less concolorous dull brown, finely, evenly, butshallowly punctured; no apparent contrasting paler markings present;hemelytral membrane bronzy-brown.Connexivum widely expanded and strikingly alternating fuscous andreddish orange, the fuscous band occupying at least the posterior three-fourths of each segment; abdominal dorsum, as seen after the wings arespread, piceous, with median sublunate orange spots on the fourth andfifth segments; hind wings dark, smoky brown.Rostrum, all pleura and legs dull reddish fuscous; sternal areas and4 NO0. 1702RUCKES: GENUS MOZENAabdominal venter dull reddish brown; a continuous dull yellow, tuber-culate ridge running diagonally across the thoracic pleura and pairs ofdull yellow, somewhat sigmoid laevigate ridges obliquely placed oneach abdominal segment. Hind femora slightly darker on the inner sur-face and provided with three longitudinal rows of very stout conicaltubercles, even in the female visible to the naked eye. Hind tibiae con-colorous throughout, even on the inner surfaces. Legs and antennae pro-vided with many fine erect setae interspersed with the tomentum.Described from one specimen.HOLOTYPE: Female, 20 mm. long; 9.0 mm. across the humeri; 10.5mm. across greatest width of abdomen. Antigua, Guatemala, August 16,1947. Collected by Dr. and Mrs. Charles Vaurie. Deposited in the Amer-ican Museum of Natural History.ALLOTYPE: None.PARATYPES: None.There appears to be a close relationship between this species and M.lanceolata (Herrich-Schaeffer), but the copius tomentose covering ofthe body readily distinguishes it along with differences in color, acute-ness of the humeri, the somewhat more widely dilated abdomen, thepiceous abdominal dorsum, and the presence of numerous fine setae. Insome respects there seems to be affinity to M. lunata (Burmeister), butthe shorter antennae and more widely expanded abdomen set it off fromthat species.Mozena pallisteri, new speciesAn elongate species, above dark fuscous to piceous, with no contrastingmarkings except a sprinkling of pale points on pronotum and hemelytra;below, variable dusky to brownish yellow, with a reddish suffusion.Head squarish as usual, fuscous above, with a slightly darker medianarea; eyes dark brown, ocelli crimson; postocular tubercles absent or atthe most obsolescent; basal three antennal segments fuscous to piceousabove, paler below, terminal segment dull orange. Antennal ratios: male:150/140/110/130; female: 140/130/100/120, i.e., segment III shortestand about three-fourths of the length of segment I.Surface of pronotum subrugose, with a scattering of small pale points;anterolateral margins before the humeri, straight with a few irregu-larly spaced, inconspicuous pale tubercles. Humeri not lunately ex-panded, protruding only a short distance beyond the base of the hemely-tra and obtusely rounded apically, with no serratulate posterior margins;diameter of pronotum across the humeri hardly, if at all, greater thanwidest diameter of the abdomen.1955 5AMERICAN MUSEUM NOVITATESScutellum obsoletely rugose, very slightly longer than wide at thebase (128 by 120), concolorous dusky to fuscous except in the femalewhere the apex is pale stramineous. Hemelytra dusky to fuscous, not con-trastingly marked except for an irregular scattering of pale points; heme-lytral membrane smoky brown. Hind wings clear, with only a very palesmoky tinge.Connexivum narrowly exposed, more or less concolorous fuscous (inthe male definitely piceous) ; if alternated, the alternations are very in-conspicuous or obsolescent. The abdominal dorsum, as seen after thewings are spread, velvety black, with a median row of at least three sub-circular luteous spots.Lateral surface of each acetabulum and the entire surface of the pro-pleuron dirty ivory; remainder of the venter variable dull brownishyellow, with a rufescent overcast. Mesosternum dusky each side of abroad median pale vitta. The oblique raised pale lines on each side ofthe abdominal segments, usually well defined in allied species, here ob-scure and somewhat diffuse. No tuberculate laevigate oblique ridges onthe thoracic pleura.Inner surfaces of all femora fuscous, outer surfaces paler and reddish;prothoracic and mesothoracic tibiae and all tarsi rufescent brown (inthe male somewhat darker); metathoracic tibiae dark fuscous to piceouson entire inner surface, outer surface dusky; in the female the hindtibiae much more dilated than in allied species provided with a largemedian spine and more nearly approaching the proportions found in themale, although the bending in the middle is less pronounced. In the maleall colors are more intensified and tend towards the darker shades.The genital segment of the male and its surrounding area contrast-ingly very dark, the cup provided with a median obtuse carina whichterminates in a short transverse ridge somewhat below the weakly con-cave apical margin of the segment.Described from three specimens.HOLOTYPE: Female, 23 mm. long; 8.25 mm. across humeri; 8 mm.wide across abdomen. Mexcala, Guerrero, Mexico, June 3, 1946. Col-lected by J. and D. Pallister. Deposited in the American Museum ofNatural History.ALLOTYPE: Male, 26 mm. long; 8.5 mm. across humeri; 8.5 mm. wideacross abdomen. Mexcala, Guerrero, Mexico, June 3, 1946. Collected byJ. and D. Pallister. Deposited in the American Museum of NaturalHistory.PARATYPE: Female. Same data as above.The very elongated form and dark color readily distinguish this species6 NO0. 1702RUCKES: GENUS MOZENAfrom all other known ones. The color approaches the black variety ofM. lunata described by Stal; this is especially true of the male. The veryinconspicuous and blunt humeri are much like those found in M. obesaMontadon, but the body proportions are again at variance. The velvetyblack abdominal dorsum with its median row of yellowish spots is quitedistinctive.Mozena lunata rufescens, new subspeciesOver-all color castaneous to reddish brown, but lighter in shade thanMozena lunata (Burmeister), of which this form apparently is a north-ern geographical race. Basal three antennal segments fuscous above,paler below, terminal one dull orange; segmental ratios: 130/120/95/120,i.e., segment III shortest, segments II and IV subequal. Postocular tuber-cles absent or obsolescent.Pronotum like that in lunata but less definitely impressed anteriorlyand less darkly colored about the humeri.Pale spots at basal angles and apex of scutellum very vague, not con-trastingly evident as in lunata. Hemelytra uniformly colored with no con-centration of pale markings on the apical border. Connexivum muchless contrastingly alternated, sometimes almost concolorous. Abdominaldorsum, as seen after the wings are spread, orange-brown to orange,with paler subcircular spots on fourth and fifth segments. In lunata theabdominal dorsum is piceous and the spotting very contrasted.The pairs of oblique ridges on each abdominal segment less raised andmore poorly defined than those in lunata, as are the diagonal tuberculatevittae on the thoracic pleura, sometimes these vittae altogether absent.Genital segments alike in both species.Described from 12 male and nine female specimens.HOLOTYPE: Male, 20 mm. long; 8 mm. across humeri; 7 mm. wideacross abdomen. Kingsville, Texas, no date. Deposited in the AmericanMuseum of Natural History.ALLOTYPE: Female, 20 mm. long; 8.25 mm. across humeri; 8 mm.wide across abdomen. Kingsville, Texas, no date. Deposited in the Ameri-can Museum of Natural History.PARATYPES: Texas: Kingsville, no date, six males, four females, de-posited in the American Museum of Natural History; Corpus Christi,July 21, 1947 (W. M. Gordon), three males, four females, deposited inthe Cornell University collection; Sutherland Springs, June 26, 1917,two males, deposited in the Cornell University collection.As stated above, this subspecies, occurring in a more northerly lati-tude, appears to be a color phase of Mozena lunata. The typical speci-1955 78 AMERICAN MUSEUM NOVITATES NO. 1702mens of lunata that I have examined come mostly from Mexico andCentral America. Only in rare instances have specimens been taken northof the Mexican border. Of almost 120 specimens of lunata in the collectionof the American Museum of Natural History, only two are recorded fromBrooks County, Texas. All others are from varying localities rangingfrom Sonora, Mexico, to Guatemala. They are invariably dark withcontrasting paler markings. In the female of the subspecies the darkand light alternations on the connexivum are more conspicuous thanthose in the male.

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