Text-to-speech devices often do a poor job of translating signals such as headings from visual into audio mode. Previous research studies have attempted to address this problem but these studies have mainly used heading detection tasks. The current study seeks to investigate 1) whether listeners find the presence of audio headings useful in natural learning tasks, and 2) the type of heading rendering that is most useful in natural learning tasks. The three learning tasks in this study include note-taking, cued recall, and knowledge transfer. Results from this study reveal that listeners find audio headings useful in the note-taking task. It is less clear how audio headings affect cued recall and knowledge transfer, but there is some evidence that a rendering strategy which conveys audio contrast plus other types of signaling information seems to facilitate cued recall performance