The brilliant appearance of our western roadsides and prairies from July to October, invites an extended study of the anatomy and physiology of the Compositae. The wide distribution and rapid increase of this family naturally call attention to dissemination and pollination. Darwin, Herman Mueller, and others, have shown at length, the direct relation between special adaptations for cross-pollination and the race stability of plants. The question now arises, what are the opportunities for cross-pollination in Compositae, and to what extent is this agent a factor on the increase and distribution of the family? No attempt is here made to go into a discussion in full of these questions for the entire Family, but simply to present a few facts relative thereto, gathered from representatives of the subtribes Heliantheae and Asterineae