This analysis confirms to a large degree the observations of Strong (1903) and Landacre (19l6). In Squalus acanthias the fifth, seventh and eighth nerves arise close together from the wall of the medulla and form a complex of roots, ganglia and fiber tracts. Taking them up in order, the gasserian or fifth nerve ganglion is seen to be a large hourglass-shaped mass extending out ventro-laterally from the brain wall: the distal part gives rise to the maxillaris V and the proximal part to the superficial opthalmic V and the mandibular V