The land snail Cochlostoma elegans (Clessin, 1879) has been found on the island of Pag (Croatia) for the first time. This is also the first ever insular finding of this species.Kopneni puž Cochlostoma elegans (Clessin, 1879) prvi puta je nađen na otoku Pagu (Hrvatska). To je ujedno prvo otočno nalazište te vrste uopće

Jasmina Mužinić

Vesna Štamol

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ISSN 1330-0520UDK 594.32(497.5/1¿13)COCHLOSTOMA ELEGANS (CLESSIN, 1879)ON THE ISLAND OF PAG (MOLLUSCA:GASTROPODA, PROSOBRANCHIA)VESNA [TAMOLDepartment of Zoology, Croatian Natural History Museum,Demetrova 1, 10000 Zagreb, CroatiaJASMINA MU@INI]Institute for Ornithology, Croatian Academy of Sciences and Arts,Ilirski trg 9, 10000 Zagreb, Croatia[tamol, V. & Mu`ini}, J.: Cochlostoma elegans (Clessin, 1879) on the island of Pag (Mol-lusca: Gastropoda, Prosobranchia). Nat. Croat., Vol. 7, No. 2, 107¿112, 1998, ZagrebThe land snail Cochlostoma elegans (Clessin, 1879) has been found on the island of Pag (Croa-tia) for the first time. This is also the first ever insular finding of this species.Key words: land snails, Cochlostoma elegans, island Pag, Croatia[tamol, V. & Mu`ini}, J.: Cochlostoma elegans (Clessin, 1879) na otoku Pagu (Mollusca:Gastropoda, Prosobranchia). Nat. Croat., Vol. 7, No. 2, 107¿112, 1998, Zagreb.Kopneni pu` Cochlostoma elegans (Clessin, 1879) prvi puta je na|en na otoku Pagu (Hrvatska).To je ujedno prvo oto~no nalazi{te te vrste uop}e.Klju~ne rije~i: kopneni pu`evi, Cochlostoma elegans, otok Pag, HrvatskaINTRODUCTIONIn Croatia, the flora and fauna of the karst caves, semi-caves and pits have notbeen extensively explored, and it thus happens that new findings may be made ofvarious kinds of cave animals and animals that rest in caves during the daytimeor at night. Examples of such insular findings are of a rare species of bat on theisland of Mljet (TVRTKOVI] & BALTI], 1996), and of a new species and genus of in-sect on the island of Cres (GIACHINO & ETONTI, 1996). Following the discovery ofNAT. CROAT. VOL. 7 No 2 107¿112 ZAGREB June 30, 1998Hrvatski prirodoslovni muzej, Demetrova 1, Zagreb, Croatiathe ferns Phyllitis scolopendrium (L.) Newm. and Ph. sagittata (DC.) Guinea et Hey-wood in the pits of the island of Pag (TRINAJSTI] & MU@INI], 1996), it was pro-posed that cave explorations should be carried out to establish the presence ofcave fauna (TRINAJSTI] & MU@INI], 1996). For this purpose, three pits were ex-plored on the island of Pag and in two of them specimens of land snails werefound.THE EXPLORED AREAThree pits from different parts of the island of Pag were explored.The island of Pag belongs to the group of the North Dalmatian islands and isseparated from the mainland by the sub-Velebit Channel. It is 63 km long, 35 kmwide, and the highest peak is Sveti Vid, 348 m. Ninety percent of its surface isrocky ground. Most of the island is made up of limestone and flysh, and groundwater running off Mount Velebit accumulates at the point where these two forma-tions meet. There are several springs, but no surface watercourses.The investigated pits are: the Bunker pit, the pit in the area of Kozar and a pitin Veli Svetojanj Bay (Fig. 1).1. The Bunker pit is located on the rocky south-western slopes at some 180metres above sea-level below the highest peak of Sveti Vid. The Stypo-Salvi-etum officinalis Horvati} (1956) 1958 plant community is predominant there(HORVATI], 1963), its main representatives being Salvia officinalis L. and Stipabromoides (L.) Brand. These slopes are a pasture ground for sheep. The pit,which has two openings 60 cm in diameter, is 1.5 m deep with a surface of3 × 1 m.2. The pit in the Veli Svetojanj Bay is located on the northern shores, 500 mwest of the ferry port, @igljen. This part of the coastline, with the SvetojanjBay was considered to be one the best localities for the construction of theferry port, but because of its specific features and the beauty of the land-scape it was not selected for this purpose. The pit has one entry point about80 cm wide and it is about 6 m deep.3. The pit in the region of Kozar is located in the northern part of the island,right above the village of Kusti}i. The pit has one opening 120 × 180 cmwide and it is 8 m deep.METHODSSnails were collected by taking individual specimens in the pits explored. Theidentification of the snails was carried out on the basis of conchological featuresas in WAGNER (1897, 1906) and WESTERLUND (1883).108 [tamol, V. &Mu`ini}, J.:Cochlostoma elegans (Clessin, 1879) on the island of PagNat. Croat. Vol. 7(2), 1998 109Fig. 1. Map of the island of Pag. The positions of the pits researched is markedby circles: 1. the »Bunker« pit; 2. the pit in the Veli Svetojanj Bay; 3. the pit inthe region of Kozar.RESULTS AND DISCUSSIONThe snail Cochlostoma elegans (Clessin, 1879) was discovered at two speleologi-cal localities on the island of Pag: the Bunker pit and the pit in the Veli SvetojanjBay. On the 14th of August 1995, some fifty live specimens of this species werenoticed on the upper surface and the lateral walls of the pit, and also on theleaves of the Phyllitis scolopendrium fern. On the 7th of August 1996, one specimenwas discovered 50 cm from the opening of the pit in the Veli Svetojanj Bay. Theentire pit in the area of Kozar was searched on the 10th of August 1996, but nosnails were found.The snails of the Cochlostoma genus are above-ground taxa. They live in a cal-careous area and feed on microflora which grows on the rocks or on dead vegeta-ble material (FECHTER & FALKNER, 1990). The discovery of the above-ground landsnail Cochlostoma elegans at two speleological sites on the island of Pag can be ac-counted for by the existence of above-ground conditions in the Bunker pit and inthe pit in Veli Svetojanj Bay. Since these are very small pits, the light, temperatureand humidity do not significantly differ from the external conditions. There hasbeen no systematic research into the above-ground or subterranean land malaco-fauna on the island of Pag, and there is no evidence of the existence of this spe-cies on Pag. It can, nevertheless, be assumed that Cochlostoma elegans is a commonspecies of the above-ground land snail fauna on this island, and that it accord-ingly also inhabits pits in which above-ground conditions largely prevail, i.e. theBunker pit and the pit in the Veli Svetojanj Bay.The genus Cochlostoma is circum-Mediterranean (KERNEY et al., 1983). In south-ern Europe it is represented by 47 species, some of which have a large number ofsubspecies (FECHTER & FALKNER, 1990). The Cochlostoma elegans species was de-scribed by CLESSIN (1879) who, as WAGNER (1897) later established, used speci-mens obtained on Mount Velebit. Soon after, numerous varieties were described(WESTERLUND, 1883; WAGNER, 1897), but their taxonomic status has so far notbeen defined. All of these varieties originate from the Velebit area. In 1906 A. J.WAGNER described the subspecies C. elegans imoschiense from the region aroundImotski and the Neretva valley, while KLEMM (1962) and SCHÜTT (1977) recordedits existence in Greece. Systematic research has been conducted into the landmalacofauna of the Adriatic islands ¿ Dugi otok (KU[^ER, 1930), Krk (BOLE, 1958),Bra~ ([TAMOL, 1986), Cres and Lo{inj ([TAMOL & VELKOVRH, 1995) ¿ but therehave been no records of the existence of the species Cochlostoma elegans. There areno records of its existence on the islands in all the literature available either, sothis can be said to be the first insular finding of Cochlostoma elegans ever made.The existence of Cochlostoma elegans on the island of Pag is the result of its con-tinuous range on the northern Croatian Adriatic coast during the last glacial stageof the Ice Age, when the island of Pag was part of the mainland. After the lastperiod of glaciation, the sea level rose and flooded the present-day Sub-VelebitChannel, making Pag an island.110 [tamol, V. &Mu`ini}, J.:Cochlostoma elegans (Clessin, 1879) on the island of PagACKNOWLEDGMENTSWe are grateful to Professor Bruno Puhari}, a speleologist and expert in the na-ture of the island of Pag, for help during the field work.Received February 23, 1998REFERENCES:BOLE, J., 1958: Zoogeografski pregled malakofavne otoka Krka. ¿ Biol. vestn. 6, 118¿123.CLESSIN, S., 1879: Zur Molluskenfauna Croatiens. ¿ Nachrichtsbl. Deutsch. Malakozool. Ges.11, 116¿125.GIACHINO, P. M., ETONTI, M., 1996: Prospelaeobates gen. nov. e due sp. n. di Leptodorinaedelle isole del Quarnero e dell’Istria (Coleoptera: Cholevidae). ¿ Acta entomol. Slov. 4(2), 63¿71.HORVATI], S., 1963: Vegetacijska karta otoka Paga s op}im pregledom vegetacijskih jedinicahrvatskog primorja. ¿ Prir. istra`. JAZU 33, Acta Biolog. 4, 1¿187.FECHTER, R., FALKNER, R., 1990: Weichtiere. pp. 287. Mosaik Verlag GmbH, München.KERNEY, M. P., CAMERON, A. D., JUNGBLUTH, J. H., 1983: Die Landschnecken Nord- und Mit-teleuropas. 384 pp. Paul Parey, Hamburg und Berlin.KLEMM, W., 1962: X. Teil. Die Gehäuseschnecken, in: BEIER, M.: Zoologische Studien inWest-Griecheland. ¿ S.-B. österr. Akad. Wiss., math.-nat. Kl. I 171, 203¿258.KU[^ER, L., 1930: Mollusca. In: VOUK, V.: Prirodoslovna istra`ivanja sjeverno-dalmatinskogoto~ja. I. Dugi i Kornati. ¿ Prir. istra`. kralj. Jugoslavije JAZU 16, 33¿39.SCHÜTT, H., 1977: Revision der griechischen Cochlostoma. ¿ Arch. Moll. 108 (1/3), 17¿35.[TAMOL, V., 1986: Prilog poznavanju kopnenih pu`eva (Gastropoda) otoka Bra~a i ugro`e-nost endema. ¿ Biosistematika 12 (1), 45¿56.[TAMOL, V., VELKOVRH, F., 1995: Contribution to the knowledge of land snail fauna of theisland of Cres and Lo{inj (Croatia) (Gastropoda: Prosobranchia, Basommatophora, Sty-lommatophora). ¿ Malak. Abh. Mus. Tierkd. Dresden 17 (20), 219¿238.TRINAJSTI], I., MU@INI], J., 1996: Phyllitis sagittata (DC.) Guinea et Heywood and Ph. scolo-pendrium (L.) Newm. (Polypodiacae) on the island of Pag (Croatia). ¿ Nat. Croat. 5 (1),83¿88.TVRTKOVI], N., BALTI], M., 1996: The Giant noctule (Nyctalus lasiopterus Schreber, 1780), firstrefinding in Croatia (Mljet island) after 69 years. ¿ Nat. Croat. 5 (1), 89¿93.WAGNER, A. J., 1897: Monographie der Gattung Pomatias Studer. ¿ Denkschr. math.- natur-wiss. Cl. kais. Akad. Wiss. 64, 565¿632, Taf. 1¿10.WAGNER, A.  J , 1906: Neue Formen und Fundorte der Genera Pomatias Stud. und AuritusWesterl. ¿ Nachrichtsb. Deutsch. malakozool. Ges. 38, 92¿101, 121¿140, Taf. 3, 4.WESTERLUND, C. A., 1883: Malakologische Miscellen II. ¿ Nachrichtsbl. Deutsch. malakozool.Ges. 15, 164¿174.Nat. Croat. Vol. 7(2), 1998 111S A @ E TA KCochlostoma elegans (Clessin, 1879) na otoku Pagu (Mollusca: Gas-tropoda, Prosobranchia)V. [tamol & J. Mu`ini}Prvi put je za otok Pag zabilje`en kopneni pu` Cochlostoma elegans koji je na-|en u kr{kim jamama Bunker i u jami u uvali Veli Svetojanj. Dosad je u Hrvat-skoj ovaj pu` poznat s podru~ja Velebita i Imotskog, u Bosni i Hercegovini izdoline Neretve u podru~ju Dre`nice, a u Gr~koj s Platanuse, pa je nalaz na otokuPagu prvi nalaz u Hrvatskoj, oto~ni i uop}e. On ukazuje na nekada{nju poveza-nost otoka Paga i susjednog kopna, to jest planine Velebita.SUMMARYCochlostoma elegans (Clessin, 1879) on the island of Pag (Mollusca:Gastropoda, Prosobranchia)V. [tamol & J. Mu`ini}The land snail Cochlostoma elegans was registered for the first time on the islandof Pag, in the karst pit Bunker and in the pit in the Veli Svetojanj Bay. Up to nowthis snail has been registered in Croatia in the Velebit and Imotski regions, in Bos-nia and Hercegovina in the Neretva river valley around Dre`nice, and in Greecein Platanusa. Thus this discovery on the island of Pag is the first insular findingof this species ever made in Croatia or elsewhere. It reflects the fact that the is-land of Pag was once part of the mainland, connected with the Velebit mountain.112 [tamol, V. &Mu`ini}, J.:Cochlostoma elegans (Clessin, 1879) on the island of Pag

Cochlostoma elegans (Clessin, 1879) na otoku Pagu (Mollusca: Gastropoda, Prosobranchia)

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ISSN 1330-0520UDK 594.32(497.5/1¿13)COCHLOSTOMA ELEGANS (CLESSIN, 1879)ON THE ISLAND OF PAG (MOLLUSCA:GASTROPODA, PROSOBRANCHIA)VESNA [TAMOLDepartment of Zoology, Croatian Natural History Museum,Demetrova 1, 10000 Zagreb, CroatiaJASMINA MU@INI]Institute for Ornithology, Croatian Academy of Sciences and Arts,Ilirski trg 9, 10000 Zagreb, Croatia[tamol, V. & Mu`ini}, J.: Cochlostoma elegans (Clessin, 1879) on the island of Pag (Mol-lusca: Gastropoda, Prosobranchia). Nat. Croat., Vol. 7, No. 2, 107¿112, 1998, ZagrebThe land snail Cochlostoma elegans (Clessin, 1879) has been found on the island of Pag (Croa-tia) for the first time. This is also the first ever insular finding of this species.Key words: land snails, Cochlostoma elegans, island Pag, Croatia[tamol, V. & Mu`ini}, J.: Cochlostoma elegans (Clessin, 1879) na otoku Pagu (Mollusca:Gastropoda, Prosobranchia). Nat. Croat., Vol. 7, No. 2, 107¿112, 1998, Zagreb.Kopneni pu` Cochlostoma elegans (Clessin, 1879) prvi puta je na|en na otoku Pagu (Hrvatska).To je ujedno prvo oto~no nalazi{te te vrste uop}e.Klju~ne rije~i: kopneni pu`evi, Cochlostoma elegans, otok Pag, HrvatskaINTRODUCTIONIn Croatia, the flora and fauna of the karst caves, semi-caves and pits have notbeen extensively explored, and it thus happens that new findings may be made ofvarious kinds of cave animals and animals that rest in caves during the daytimeor at night. Examples of such insular findings are of a rare species of bat on theisland of Mljet (TVRTKOVI] & BALTI], 1996), and of a new species and genus of in-sect on the island of Cres (GIACHINO & ETONTI, 1996). Following the discovery ofNAT. CROAT. VOL. 7 No 2 107¿112 ZAGREB June 30, 1998Hrvatski prirodoslovni muzej, Demetrova 1, Zagreb, Croatiathe ferns Phyllitis scolopendrium (L.) Newm. and Ph. sagittata (DC.) Guinea et Hey-wood in the pits of the island of Pag (TRINAJSTI] & MU@INI], 1996), it was pro-posed that cave explorations should be carried out to establish the presence ofcave fauna (TRINAJSTI] & MU@INI], 1996). For this purpose, three pits were ex-plored on the island of Pag and in two of them specimens of land snails werefound.THE EXPLORED AREAThree pits from different parts of the island of Pag were explored.The island of Pag belongs to the group of the North Dalmatian islands and isseparated from the mainland by the sub-Velebit Channel. It is 63 km long, 35 kmwide, and the highest peak is Sveti Vid, 348 m. Ninety percent of its surface isrocky ground. Most of the island is made up of limestone and flysh, and groundwater running off Mount Velebit accumulates at the point where these two forma-tions meet. There are several springs, but no surface watercourses.The investigated pits are: the Bunker pit, the pit in the area of Kozar and a pitin Veli Svetojanj Bay (Fig. 1).1. The Bunker pit is located on the rocky south-western slopes at some 180metres above sea-level below the highest peak of Sveti Vid. The Stypo-Salvi-etum officinalis Horvati} (1956) 1958 plant community is predominant there(HORVATI], 1963), its main representatives being Salvia officinalis L. and Stipabromoides (L.) Brand. These slopes are a pasture ground for sheep. The pit,which has two openings 60 cm in diameter, is 1.5 m deep with a surface of3 × 1 m.2. The pit in the Veli Svetojanj Bay is located on the northern shores, 500 mwest of the ferry port, @igljen. This part of the coastline, with the SvetojanjBay was considered to be one the best localities for the construction of theferry port, but because of its specific features and the beauty of the land-scape it was not selected for this purpose. The pit has one entry point about80 cm wide and it is about 6 m deep.3. The pit in the region of Kozar is located in the northern part of the island,right above the village of Kusti}i. The pit has one opening 120 × 180 cmwide and it is 8 m deep.METHODSSnails were collected by taking individual specimens in the pits explored. Theidentification of the snails was carried out on the basis of conchological featuresas in WAGNER (1897, 1906) and WESTERLUND (1883).108 [tamol, V. & Mu`ini}, J.: Cochlostoma elegans (Clessin, 1879) on the island of PagNat. Croat. Vol. 7(2), 1998 109Fig. 1. Map of the island of Pag. The positions of the pits researched is markedby circles: 1. the »Bunker« pit; 2. the pit in the Veli Svetojanj Bay; 3. the pit inthe region of Kozar.RESULTS AND DISCUSSIONThe snail Cochlostoma elegans (Clessin, 1879) was discovered at two speleologi-cal localities on the island of Pag: the Bunker pit and the pit in the Veli SvetojanjBay. On the 14th of August 1995, some fifty live specimens of this species werenoticed on the upper surface and the lateral walls of the pit, and also on theleaves of the Phyllitis scolopendrium fern. On the 7th of August 1996, one specimenwas discovered 50 cm from the opening of the pit in the Veli Svetojanj Bay. Theentire pit in the area of Kozar was searched on the 10th of August 1996, but nosnails were found.The snails of the Cochlostoma genus are above-ground taxa. They live in a cal-careous area and feed on microflora which grows on the rocks or on dead vegeta-ble material (FECHTER & FALKNER, 1990). The discovery of the above-ground landsnail Cochlostoma elegans at two speleological sites on the island of Pag can be ac-counted for by the existence of above-ground conditions in the Bunker pit and inthe pit in Veli Svetojanj Bay. Since these are very small pits, the light, temperatureand humidity do not significantly differ from the external conditions. There hasbeen no systematic research into the above-ground or subterranean land malaco-fauna on the island of Pag, and there is no evidence of the existence of this spe-cies on Pag. It can, nevertheless, be assumed that Cochlostoma elegans is a commonspecies of the above-ground land snail fauna on this island, and that it accord-ingly also inhabits pits in which above-ground conditions largely prevail, i.e. theBunker pit and the pit in the Veli Svetojanj Bay.The genus Cochlostoma is circum-Mediterranean (KERNEY et al., 1983). In south-ern Europe it is represented by 47 species, some of which have a large number ofsubspecies (FECHTER & FALKNER, 1990). The Cochlostoma elegans species was de-scribed by CLESSIN (1879) who, as WAGNER (1897) later established, used speci-mens obtained on Mount Velebit. Soon after, numerous varieties were described(WESTERLUND, 1883; WAGNER, 1897), but their taxonomic status has so far notbeen defined. All of these varieties originate from the Velebit area. In 1906 A. J.WAGNER described the subspecies C. elegans imoschiense from the region aroundImotski and the Neretva valley, while KLEMM (1962) and SCHÜTT (1977) recordedits existence in Greece. Systematic research has been conducted into the landmalacofauna of the Adriatic islands ¿ Dugi otok (KU[^ER, 1930), Krk (BOLE, 1958),Bra~ ([TAMOL, 1986), Cres and Lo{inj ([TAMOL & VELKOVRH, 1995) ¿ but therehave been no records of the existence of the species Cochlostoma elegans. There areno records of its existence on the islands in all the literature available either, sothis can be said to be the first insular finding of Cochlostoma elegans ever made.The existence of Cochlostoma elegans on the island of Pag is the result of its con-tinuous range on the northern Croatian Adriatic coast during the last glacial stageof the Ice Age, when the island of Pag was part of the mainland. After the lastperiod of glaciation, the sea level rose and flooded the present-day Sub-VelebitChannel, making Pag an island.110 [tamol, V. & Mu`ini}, J.: Cochlostoma elegans (Clessin, 1879) on the island of PagACKNOWLEDGMENTSWe are grateful to Professor Bruno Puhari}, a speleologist and expert in the na-ture of the island of Pag, for help during the field work.Received February 23, 1998REFERENCES:BOLE, J., 1958: Zoogeografski pregled malakofavne otoka Krka. ¿ Biol. vestn. 6, 118¿123.CLESSIN, S., 1879: Zur Molluskenfauna Croatiens. ¿ Nachrichtsbl. Deutsch. Malakozool. Ges.11, 116¿125.GIACHINO, P. M., ETONTI, M., 1996: Prospelaeobates gen. nov. e due sp. n. di Leptodorinaedelle isole del Quarnero e dell’Istria (Coleoptera: Cholevidae). ¿ Acta entomol. Slov. 4(2), 63¿71.HORVATI], S., 1963: Vegetacijska karta otoka Paga s op}im pregledom vegetacijskih jedinicahrvatskog primorja. ¿ Prir. istra`. JAZU 33, Acta Biolog. 4, 1¿187.FECHTER, R., FALKNER, R., 1990: Weichtiere. pp. 287. Mosaik Verlag GmbH, München.KERNEY, M. P., CAMERON, A. D., JUNGBLUTH, J. H., 1983: Die Landschnecken Nord- und Mit-teleuropas. 384 pp. Paul Parey, Hamburg und Berlin.KLEMM, W., 1962: X. Teil. Die Gehäuseschnecken, in: BEIER, M.: Zoologische Studien inWest-Griecheland. ¿ S.-B. österr. Akad. Wiss., math.-nat. Kl. I 171, 203¿258.KU[^ER, L., 1930: Mollusca. In: VOUK, V.: Prirodoslovna istra`ivanja sjeverno-dalmatinskogoto~ja. I. Dugi i Kornati. ¿ Prir. istra`. kralj. Jugoslavije JAZU 16, 33¿39.SCHÜTT, H., 1977: Revision der griechischen Cochlostoma. ¿ Arch. Moll. 108 (1/3), 17¿35.[TAMOL, V., 1986: Prilog poznavanju kopnenih pu`eva (Gastropoda) otoka Bra~a i ugro`e-nost endema. ¿ Biosistematika 12 (1), 45¿56.[TAMOL, V., VELKOVRH, F., 1995: Contribution to the knowledge of land snail fauna of theisland of Cres and Lo{inj (Croatia) (Gastropoda: Prosobranchia, Basommatophora, Sty-lommatophora). ¿ Malak. Abh. Mus. Tierkd. Dresden 17 (20), 219¿238.TRINAJSTI], I., MU@INI], J., 1996: Phyllitis sagittata (DC.) Guinea et Heywood and Ph. scolo-pendrium (L.) Newm. (Polypodiacae) on the island of Pag (Croatia). ¿ Nat. Croat. 5 (1),83¿88.TVRTKOVI], N., BALTI], M., 1996: The Giant noctule (Nyctalus lasiopterus Schreber, 1780), firstrefinding in Croatia (Mljet island) after 69 years. ¿ Nat. Croat. 5 (1), 89¿93.WAGNER, A. J., 1897: Monographie der Gattung Pomatias Studer. ¿ Denkschr. math.- natur-wiss. Cl. kais. Akad. Wiss. 64, 565¿632, Taf. 1¿10.WAGNER, A. J , 1906: Neue Formen und Fundorte der Genera Pomatias Stud. und AuritusWesterl. ¿ Nachrichtsb. Deutsch. malakozool. Ges. 38, 92¿101, 121¿140, Taf. 3, 4.WESTERLUND, C. A., 1883: Malakologische Miscellen II. ¿ Nachrichtsbl. Deutsch. malakozool.Ges. 15, 164¿174.Nat. Croat. Vol. 7(2), 1998 111S A @ E TA KCochlostoma elegans (Clessin, 1879) na otoku Pagu (Mollusca: Gas-tropoda, Prosobranchia)V. [tamol & J. Mu`ini}Prvi put je za otok Pag zabilje`en kopneni pu` Cochlostoma elegans koji je na-|en u kr{kim jamama Bunker i u jami u uvali Veli Svetojanj. Dosad je u Hrvat-skoj ovaj pu` poznat s podru~ja Velebita i Imotskog, u Bosni i Hercegovini izdoline Neretve u podru~ju Dre`nice, a u Gr~koj s Platanuse, pa je nalaz na otokuPagu prvi nalaz u Hrvatskoj, oto~ni i uop}e. On ukazuje na nekada{nju poveza-nost otoka Paga i susjednog kopna, to jest planine Velebita.SUMMARYCochlostoma elegans (Clessin, 1879) on the island of Pag (Mollusca:Gastropoda, Prosobranchia)V. [tamol & J. Mu`ini}The land snail Cochlostoma elegans was registered for the first time on the islandof Pag, in the karst pit Bunker and in the pit in the Veli Svetojanj Bay. Up to nowthis snail has been registered in Croatia in the Velebit and Imotski regions, in Bos-nia and Hercegovina in the Neretva river valley around Dre`nice, and in Greecein Platanusa. Thus this discovery on the island of Pag is the first insular findingof this species ever made in Croatia or elsewhere. It reflects the fact that the is-land of Pag was once part of the mainland, connected with the Velebit mountain.112 [tamol, V. & Mu`ini}, J.: Cochlostoma elegans (Clessin, 1879) on the island of Pag

Cochlostoma elegans (Clessin, 1879) na otoku Pagu (Mollusca: Gastropoda, Prosobranchia)

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