0. A. Sæthern

L. C. . Ferrington Jr

CHIRONOMUS Journal of Chironomidae Research

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14CURRENT RESEARCHNEW COMBINATIONS AND SYNONYMS IN EUROPEANPseudosmittia GOETGHEBUER AND RELATED GENERA.By 0. A. Sæther & L. C. Ferrington Jr.Museum of Zoology. University of Bergen, Bergen, Norwayand Department of Entomology University of Minnesota, St. Paul, Minnesota. USAWhen one of us (0. A. S) was on sabbaticalwith the other at the University of Kansas arevision of the genus PseudosmittiaGoetghebuer was started and a preliminarymanuscript describing and re-describing nearly100 species assignable to the genus wascompleted. Due to various circumstances,including a change of venue for one of us (L.C. F) from Lawrence, Kansas, to theUniversity of Minnesota in St. Paul,Minnesota, the publication date has beendelayed. Recently, however, one of us (0. A.S.) was requested to summarize thechironomids in the Fauna Europaea project, adatabase which will include all terrestrial andaquatic animals of Europe with theirdistribution in respective countries. Furtherinformation is obtainable onhttp://www.faunaeur.org. The database willeventually result in a variety of publicationsfor the different groups, but new synonyms andcombinations known to us should not beintroduced in the data base, and we believe it isnot desirable to use knowingly wrong namesand combinations in the data base. We thusfind it necessary to publish a list of the typeswhich will cause changes to the nomenclatureof the European species. We have examinedthe type material of most relevant museums,but a few remain to be investigated. Here weare listing only new combinations, newsynonyms, new nomina dubia or other changesrelative to the catalogue by Ashe & Cranston(1990).The following abbreviations for collections areused:BMNH: The Natural History Museum (BritishMuseum, Natural History), London. EnglandIRSN: Institute Royal de Science Naturelles deBelgique, Bruxelles, Belgium.NMS: Natur-Museum Senckenherg, FrankfurtAm Main, Germany.ZMB: Museum of Zoology, University ofBergen, Norway.ZMH: Zoological Museum, University ofHelsinki, Finland.ZMO: Zoological Museum. University ofOslo, Norway.ZSM: Zoologisches Staatssammlung, Munich,Germany.New generic placementsDactylocladius albipennis Goetghebuer, 1921:85[= Pseudosmittia albipennis (Goetghebuer)comb. n.] Holotype male in IRSN.Orthocladius (Dactylocladius) brevifurcatusEdwards, 1926: 781[= Pseudosmittia brevifurcata (Edwards)comb. n.] Types in BMNH.Lindebergia bothnica Tuiskunen,  1984:121[= Pseudosmittia bothnica (Tuiskunen) comb.n.] Holotype male in ZMH.Smittia lacunarum Goetghebuer,  1931: 217[= Bryophaenocladius lacunarum(Goetghebuer) comb. n.] Holotype male inIRSN.Smittia (Pseudosmittia) terrestris Goetghebuer,1943: 109[= Bryophaenocladius terrestris (Goetghebuer)comb. n.] Holotype female in IRSN.New synonymsPseudosmittia amamibifurca Sasa, 1990: 132(=Pseudosmittia mathildae Albu, 1968, syn.n.)Pseudosmittia antillaria Sæther,  1981: 29 [=Pseudosmittia forcipata (Goetghebuer, 1921)syn. n.]  Holotype male in ZMB.Pseudosmittia arenaria flava Strenzke, 1960:419 (= Pseudosmittia arenaria Strenzke, 1960,syn. n.) Holotype female in NMS.Smittia avicularia Goetghebuer,  1950: 13 [=Pseudosmittia trilobata (Edwards, 1929) syn.n.] Holotype male in IRSN.15Smittia brachyptera Goetghebuer, 1934: 389[= Pseudosmittia conjuncta (Edwards, 1929)comb. n.] Holotype male in IRSN.Pseudosmittia brevitarsis Brundin, 1947: 40 (=Pseudosmittia ruttneri Strenzke & Thienemann1942, syn. n.). The type could not be located.Spaniotoma (Smittia) curticosta Edwards,1929: 364 [= Pseudosmittia albipennis(Goetghebuer, 1921) syn. n.] Holotype male inBMNH.Smittia hamata Freeman, 1956: 355 [=Pseudosmittia danconai (Marcuzzi, 1947) syn.n.] Holotype male in NMS. Paratypes inBMNH also examined.Pseudosmittia linguata Caspers & Reiss, 1989:128 (= Pseudosmittia nishiharaensis Sasa &Hasegawa, 1988: 247). Holotype male in ZSM.Pseudosmittia kurobaokasia Sasa & Okazawa,1992a: 57 (= Pseudosmittia ruttneri Strenzke& Thienemann, 1942. syn. n.)Smittia longitibia Goetghebuer, 1933: 29 [=Pseudosmittia nanseni (Kieffer, 1926) syn. n.]Holotype male in ZMO.Pseudosmittia mediocarinata Caspers & Reiss,1989: 132 [= Pseudosmittia nanseni (Kieffer1926) syn. n.] Holotype male in ZSM.Smittia oxoniana Edwards, 1937: 146 notEdwards, 1922:204 (= Pseudosmittia ruttneriStrenzke & Thienemann, 1942  syn. n.)Spaniotoma (Smittia) recta Edwards, 1929:362 [= Pseudosmittia oxoniana (Edwards),Edwards, 1922, not Edwards, 1937, syn. n.]Pseudosmittia schachti Caspers & Reiss,1989: 130 (= Pseudosmittia ruttneri Strenzke& Thienemann, 1942, syn. n.) Holotype malein ZSM. Paratype male misidentified P.oxoniana (Edwards).Spaniotoma (Smittia) scotica Edwards, 1929:363 [= Camptocladius stercorarius (De Geer,1776) syn. n.] Holotype female in BMNH.Smittia (Orthosmittia) subrecta Goetghebuer,1942: 112 [= Parakieffèriella coronata(Edwards, 1929) syn. n.] Types in IRSN.Pseudosmittia togadistalis Sasa, Watanabe &Arakawa,1992: 233 [=Pseudosmittia gracilis(Goetghebuer, 1913) syn. n.]Pseudosmittia togarisea Sasa & Okazawa,1992b: 160 [=Pseudosmittia oxoniana(Edwards, 1922) syn. n.]Pseudosmittia togasitea Sasa & Okazawa,1992b: 161 [=Camptocladius stercorarius (DeGeer, 1776) syn. n.]Pseudosmittia togativea Sasa & Okazawa,1992b: 162 [=Camptocladius stercorarius (DeGeer, 1776) syn. n.]Smittia triappendiculata Goetghebuer, 1931:216 [= Pseudosmittia forcipata (Goetghebuer,1921) syn. n.] Holotype male in IRSN.Negation of synonymyPseudosmittia triplex Strenzke, 1950: 301.Valid species and not a synonym of P.forcipata (Goetghebuer, 1921) as stated inAshe & Cranston (1990). Holotype male inNMS.New statusPseudosmittia virgo montana Strenzke, 1950:303 (= Pseudosmittia montana Strenzke stat.n.) Holotype male with pupal exuviae in NMS.Nomina dubiaCamptocladius flaviventris Kieffer, 192la: 289,nomen dubium. The type could not be located.Camptocladius hexalobus Kieffer, 1924: 395(= Pseudosmittia hexalobus Kieffer, nomendubium). Type lost. A likely senior synonymof either P. trilobata or P. obtusa.Camptocladius longicrus Kieffer, 1921b: 100,nomen dubium. Type lost. A pupal exuviaefrom NMS is marked type, but collected in1941 and thus wrongly marked.Pseudosmittia restricta Brundin, 1956: 170,nomen dubium. The type could not be found.The species cannot belong in Pseudosmittia ifthe description is correct as a distinct scutalhump is present and the genitalia differ fromall other known species.Smittia (Pseudosmittia) tenebrosaGoetghebuer, 1943: 109 [= Pseudosmittiatenebrosa (Goetghebuer) nomen dubium].Holotype male in IRSN. Mounted between twoplastic strips on a pin and ruined in attempt ofremounting. Probably a synonym of P. simplexStrenzke & Thienemann.Camptocladius trifoliatus Kieffer, 1924: 73,nomen dubium. The type could not be located.Smittia (Pseudosmittia) vicana Goetghebuer,1943: 110 [= Pseudosmittia vicana(Goetghebuer), nomen dubium]. The femaleholotype supposedly in IRSN could not befound.16ReferencesALBU. P. 1968. Pseudosmittia mathildae sp. n.(Diptera, Chironomidae). - Annls Zool. fenn. 5:4-5.ASHE, P. & CRANSTON. P. S. 1990. FamilyChironomidae. Pages 113-355. in Soos A. andPAPP L. (eds): Catalogue of Palaearctic Diptera.Vol. 2 Psychodidae — Chironomidae. -Akadémia Kiadó, Budapest, 499 pp.BRUNDIN. L. 1947. Zur Kenntnis der schwedischenChironomiden. - Ark. Zool. 39 A: 1-95.BRUNDIN, L. 1956. Zur Systematik derOrthocladiinae (Dipt., Chironomidae). -Rep.Inst.Freshwat. Res. Drottningholm 37: 5-185.CASPERS. N. & REISS. F. 1989. Die Chironomidaeder Türkei. Teil 1: Podonominae, Diamesinae,Prodiamesinae, Orthocladiinae (Diptera,Nematocera. Chironomidae). - Entomofauna 10:105—160.EDWARDS, F. W. 1922. Results of the OxfordUniversity Expedition to Spitsbergen, 1921. -Ann. Mag. Nat. Hist. (9) 10: 193-215.EDWARDS, F. W. 1926. On marine Chironomidae(Diptera) with descriptions of a new genus andfour new species from Samoa. - Proc. Zool.Soc. Lond., 1926: 779-806.EDWARDS. F. W. 1929. British non-biting midges(Diptera, Chironomidae). - Trans. Ent. Soc.Lond. 77: 279-430.EDWARDS, F. W. 1937. Chironomidae (Diptera)collected by Prof. Aug. Thienemann in SwedishLappland. - Ann. Mag. Nat. Hist. (10) 20: 140-148.GOETGHEBUER, M. 1921. Chironomides deBelgique et spécialement de la zone desFlandres. - Mém. Mus. R. Hist. Nat. Belg. 8:1-211.GOETGHEBUER, M. 1931. Ceratopogonidae etChironomidae nouveaux d’Europe. - Bull. AnnlsSoc. R. ent. Belg. 71: 211-218.GOETGHEBUER, M. 1933. Chironomides duGroenland oriental, du Svalbard et de la terre deFrancois Joseph. - Skr. Svalbard Ishavet 53: 19-31.0GOETGHEBUER, M. 1934. Note sur un nouveauChironomide brachyptêre. - Bull. Annls Soc. R.ent. Belg. 74: 388-390.GOETGHEBUER, M. 1942. 13g. Tendipedidae(Chironomidae). f) Subfamilie Orthocladiinae.A. Die Imagines (part). - Fliegen pal. Reg. 3(1):25-64.GOETGHEBUER, M. 1950. Ceratopogonidae etChironomidae nouveaux ou peu connusd’Europe (Treizième note). - Bull. Inst. R. Sci.nat. Belg. 25 (14): 1-8.KIEFFER, J. J. 1921a. Chironomides de Courlande. -Annls Soc. Scient. Brux. 40: 275-298.KIEFFER, J. J. 1921b. Chironomides nouveaux oupeu connus de la region paléarctique. - Bull. Soc.Hist. Nat. Metz 29: 51-109.KIEFFER, J. J. 1924. Quelques Chironomidesnouveaux et remarquables du Nord de 1’Europe.- Annls Soc. Scient. Brux 43: 390-397.KIEFFER, J. J. 1926. Chironomiden der 2. Fram-Expedition (1898-1902). - Norsk ent. Tidsskr.(1925) 2: 78-89.MARCUZZI, G. 1947. Descrizione di tre nuovespecie di Smittia della Laguna di Venezia. - Boll.Soc. Ent. ital. 77: 9-13.PINDER, L. C. V. 1978. A key to adult males ofBritish Chironomidae. Part 1. The key; part 2.Illustrations of the hypopygia. - Scient. PublsFreshwat. biol. Ass. 37, 169 pp.; 189 Figs.SÆTHER, 0. A. 1981. Orthocladiinae (Diptera.Chironomidae) from the British West Indies,withdescriptions of Antillocladius n. gen.,Lipurometriocnemus n. gen. Compterosmittian.gen. and Diplosmittia n. gen. - Ent. scand.,Suppl. 16, 46 pp.SASA, M. 1984. Studies on chironomid midges inlakes of the Nikko National Park. Pt. II.Taxonomical and morphological studies on thechironomid species collected from lakes in theNikko National Park.  - Res. Rep. Nat. inst.envir. Stud. 70: 16-2 15.SASA, M. 1990. Studies on the chironomid midges(Diptera, Chironomidae) of the Nansei Islands,Southern Japan.  - Japan J. Exp. Med. 60: 111-165.SASA, M.. & HASEGAWA, H. 1988. Additionalrecords of the chironomid midges from theRyukyu Islands, southern Japan (Diptera,Chironomidae). - Jpn. J. sanit. Zool. 39: 229-256.SASA, M.. & OKAZAWA, T. 1992a. Studies on thechironomid midges (yusurika) of Kurobe River. -Res. Rep. Toyama Pref. Envir. Poll. Res. Centre1992: 40-91.SASA, M. & OKAZAWA, T. 1992 b. Studies on thechironomid midges (yusurika) of Toga-Mura.Toyama. Part 2. The subfamily Orthocladiinae.-Res. Rep. Toyama Pref. Envir. Poll. Res. Centre1992: 92-204.SASA, M., WATANABE M. & ARAKAWA, R. 1992.Additional records of Chironomidae from Toga-Mura, 1992. - Res. Rep. Toyama Pref. Envir.Poll. Res. Centre 1992: 231- 246.STRENZKE, K. 1950. Systematik. Morphologie undOkologie der terrestrischen Chironomiden. -Arch. Hydrobiol. Suppl. 18: 207-414.STRENZKE, K. & THIENEMANN, A. 1942. Zwei neuePseudosmittia-Arten (Diptera, Chironomidae)aus dem Gebiete der Lunzer Seen. - Int. Revueges. Hydrobiol. Hydrogr. 42: 356-387.TUISKUNEN, J. 1984. Lindebergia bothnica. gen. n.,sp. n. 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NEW COMBINATIONS AND SYNONYMS IN EUROPEAN Pseudosmittia GOETGHEBUER AND RELATED GENERA

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NEW COMBINATIONS AND SYNONYMS IN EUROPEAN Pseudosmittia GOETGHEBUER AND RELATED GENERA

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