Permian Chonetoidea and Spiriferoidea of Australasia : Gondwanan relationships, provincialism, palaeobiogeography

Abstract

Members of the Spiriferoidea are dominant in the Permian marine invertebrate faunas of Australia. Genera can be endemic to a particular province, demonstrate a wider Gondwanan and peri-Gondwanan distribution and several indicate a bipolar distribution. Australasian spiriferoids are included within the families Spiriferidae, Neospiriferidae, Trigonotretidae and the Spiriferellidae. Several genera and species are the largest spirferoids ever recorded. Few genera are shared between the Westralian and Austrazean provinces but a higher proportion of genera are shared between the Westralian province and the Cimmerian Realm.Representatives of the Chonetoidea are a less common but significant element of the faunas. They were used over 20 years ago to define the Westralian and Austrazean provinces of Australasia &ndash; concepts that are in widespread use today. The Paratinan and Cimmerian provinces were also defined at that time despite difficulties in their definition. Through more recent studies the Cimmerian Province has been upgraded to a Realm while the Paratinan Province is more clearly defined for the earliest Permian, based on the chonetoids of Patagonia and western and eastern central Argentina.Distribution of the various genera is best explained by an interplay of factors including surface and deeper oceanic currents, marine water temperatures and tectonic events such as the clockwise rotation of Gondwanan and the dispersal of the peri-Gondwanan Cimmerian terranes. Austrazean faunas developed in isolation under the influence of cooler and cold waters during the early Permian. Late Permian faunas demonstrated more widespread linkages.<br /