Ibaragi Yasushi

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Miscanthidium junceum subsp. teretifolium （Poaceae）; a new combination for an Africangrass著者 Ibaragi Yasushijournal orpublication title植物地理・分類研究 = The journal ofphytogeography and toxonomyvolume 52number 2page range 175-178year 2004-12-30URL http://hdl.handle.net/2297/48798Miscanthidium teretifolium（Stapf）Stapf hasoften been treated synonymously with M .junceum（Stapf）Stapf, since diagnostic character-istics are quantitative and vary（Launert 1970 ;Gibbs Russell et al. 1985 ; Gibbs Russell 1991 ;Clayton 2002）. However they differ in the vari-ation of callus and glume hair（Fig. 1）and geo-graphic distribution（Fig.2）. Additionally, severaldifferences are observed in the inflorescencelength and awn length（Stapf 1906 ; Pilger1907 ; Chippindall 1955）. Therefore, it is appro-priate to classify them at subspecific rank in M .junceum.Miscanthidium Stapf was treated as an inde-pendent genus here as it differs from the genusMiscanthus Andersson, by chromosome number（De Wet 1960 ; Hoshino and Davidse 1988）andphylogeny（Hodkinson et al. 1997, 2002）.Miscanthidium junceum（Stapf）Stapf in Hooker’sIcon. Pl. 31 : t. 3084（1922）- Chippindall inMeredith, Grasses and Pastures S. Africa : 480（1955）- Launert in Merxmüller, Prodr. Fl. SW.Africa 160 : 131（1970）- Erianthus junceus Stapfin Dyer, Fl. Cap. 7 : 324（1898）- Miscanthusjunceus（Stapf）Pilg. in Nat. Pflanzenfam.（ed.2）14 e : 113（1940）- Gibbs Russell in Grasses ofSouthern Africa : 221（1991）. Types : Kalahariregion. Basutoland. Leribe, Buchanan 228 ;Transvaal. Olifants river, Nelson 73 ; Junglestreamlet, Nelson 77（Syntypes : WU? n.v. ）Key to the subspecies of Miscanthidium junceum1. Culm up to 1-2.5 m high ; inflorescence 25-42cm long ; glume hairs 1.5-3 mm long, golden ;callus hairs 1-2.5 mm long ; awn 3-6 mm long---------------------------------------------subsp. junceum1. Culm up to 2-3.5 m high ; inflorescence 32-51cm long ; glume hairs 3-6.5 mm long, silver ;callus hairs 1-3 mm long ; awn 6-10 mm long----------------------------------------subsp. teretifoliumMiscanthidium junceum subsp. junceumCleistachne teretifolia Hack. in Oesterr. Bot. Z.51 : 153（1901）- Miscanthidium gossweileri Stapfin Prain, Fl. Trop. Afr. 9 : 90（1917）- Miscanthusgossweileri（Stapf）Pilg. in Nat. Pflanzenfam.（ed.2）14 e : 113（1940）. Type : Angola. Campos deHumpata prope rivum Nene. Newton a. 1883（Holotype : W?. n.v.）.Distribution : Angola, Zambia, Zimbabwe, SouthAfrica and Swaziland.Specimens examined : without precise locality,Zeyher 1828（BM）.Journal of Phytogeography and Taxonomy 52 : 175-178, 2004The Society for the Study of Phytogeography and Taxonomy 2004Yasushi Ibaragi : Miscanthidium junceum subsp. teretifolium（Poaceae）; a new combination for an African grassTokushima Prefectural Museum, Bunka-no-mori, Hachiman-cho, Tokushima 770―8070, JapanFig. 1. Comparison of the maximum length of thecallus hairs and the glume hairs of Miscanthidiumjunceum subsp. junceum（○）and subsp. teretifolium（●）. Dots are based on all the specimens examinedin this study.－ 175 －Angola. Benguella : Tomens dusky purple inthe maarslus of the rivulet Luasenha-lutchi,Gossweiler 3067（BM）；Moxico : R.Luena, VilaLuso, alt. ca. 1,240 m, Exell & Mendonca 1546（BM） ; along the river Tiengs Kuito, Gossweiler4032（BM）.Zambia. Barotseland : Mankoya, Luena riverbank Mangango Mission, Verboom 1186（BM）.Zimbabwe. Victoria Falls, S. bank of Zambesi,alt. 3,000 ft, Rogers 5664（BM, BOL）; SalisburySouth, 25 Jan. 1920, Eyles 2032（BM）; SalisburyEast, Eyles 1553（BM）.South Africa. Natal : Margate near airfield,alt. 75 m, Ellis 6020（PRE） ; Utrecht, alt. ca.1,550―1,650 m, Devenish 1736（PRE） ; 39 km S.of Utrecht, along road to Blood River Monument,Davids 6822（PRE）; Transvaal : near Potgieters-rust, alt. 5,500 ft, Galpin 8878（PRE） ; 10 m E.of Ylammanskraal, Winter & Codd 251（PRE） ;Lochiel, alt. 1,500 m, Werdermann & Oberdieck2224（PRE）; Lydenburg, bei der Stadt Lydnburg,Wilms 1896（BM） ; Melspncut, White River, alt.3,000 ft, Winter & Codd 444（BM） ; Pietersburg,north of Naboomspruit Farm, Kaalfontein, alt.1,600 m, Burgoyne 1739（PRE） ; Potchefstroom,Louw 1633（BM） ; Potchefstroom, Louw 1466（BM） ; Rustenburg, Bartlett’s Farm, Magalies-berg Mountains, alt. 1,372 m, Mogg 36178（PRE）; Sibasa, Tate Vondo, alt. 1,178 m, Hemm559（PRE）; near Pretoria, Crusoos 11138（BM）;Vaalwater, Emaweni Game Lodge, Waterval Ma-couwkuil navy water, Theron 259（PRE） ; about9 miles west of Krugersdorp, on Farm Gladys-vale, Rodin 3943（NBG）; Thabazimbi KransbergKloof on plateau, alt. 1,400 m, Westfall 1006（PRE） ; Ohrigstaddam, Nat. Res., alt. 4,600 ft,Jacobsen 2641（PRE）; Bronkhorstspruit, Roode-poort 10 km west of town, alt. 1,402 m, Turner18（PRE） ; alt. 3,500 ft, Schlechter 6903（BM） ;Lebowa, alt. 2650 ft, Ellery 382（PRE）.Swaziland. Hhohho, Malolotjya Nature Re-serve, stream east of staff house, just below con-fluence of stream from east and stream belowroad, Braun 1363（PRE） ; Hlatikuen, SiboweFig. 2. Distribution map of Miscanthidium junceum subsp. junceum and subsp. teretifolium.植物地理・分類研究 第 52巻第 2号 2004年 12月－ 176 －River, alt. ca. 2,000 ft, Compton 29706（NBG） ;Manhaiana, alt. ca. 3,000 ft, Compton 32230（NBG）; Mbabane, Hlambanyati valley, Compton24947（NBG） ; Mbabane, Black Mhluzi Cause-way, alt. ca. 3,500 ft, Compton 30428（NBG）.Miscanthidium junceum subsp. teretifolium（Stapf）Ibaragi, comb. et stat. nov.Erianthus teretifolius Stapf in J. Linn. Soc.Bot. 37 : 478（1906）- Miscanthidium teretifolium（Stapf）Stapf in Prain, Fl. Trop. Afr. 9 : 89（1917）- Stapf in Hooker’s Icon. Pl. 31 : t. 3084（1922）- Chippindall in Meredith, Grasses andPastures S. Africa : 480（1955）- Miscanthusteretifolius（Stapf）Pilg. in Nat. Pflanzenfam.（ed.2）14 e : 113（1940）. Type : Zimbabwe. VictoriaFalls, banks of river and islands. Fl. and Fr. Sep.1905, Gibbs 141（Holotype : K, n.v. ; Isotype :BM）.Erianthus teretifolius Pilg. in Bot. Jahrb. Syst.39 : 597（1907）non Stapf（1906）. Type : Angola.Malange, Quisanya, near Panca Meueba,Gossweiler 807（Holotype : BM）.Distribution : Angola, Zambia, Zimbabwe, Na-mibia and Botswana.Additional specimens examined : Angola.Santa Cruz, Mashi Rivel, near Shangombo, alt.3,400 ft, Codd 7498（BM）.Zambia. Barotseland : 8 miles N. of Nang-weshi, alt. 3,400 ft, Codd 7163（BM） ; Namwala,Kafue National Park, Mitchell 14/82（BM）.Zimbabwe. Gokwe, Gokwe Lly Gate，±10 m,N.W. of Gokwe, Baisgham 637（BM）.Namibia. Andara, R.C. Miss. Station camp,Müller 518（PRE）.Botswana. Becl Avaland, Sherobe, S. van Son28607（BM） ; Northern, Khwai River, 1/2 hoursboat trip upstream from Gobegha lagoon,Okavango swamp, alt. 1,000 m, Gibbs Russell &Biegel 1546（PRE）; Northern, Mboma, Okavangoswamps, Smith 91（PRE） ; Northern, DobongaIsland, Smith 656（PRE） ; Okavango, imOmurmmba , Omatake Bai , Blackfontein ,nordlich Bumbi, Giess 10096（PRE）.AcknowledgementsI wish to express my thanks to the curatorsand staffs of BM, BOL, NBG, PRE and SAM forproviding specimens. I am also indebted to Dr.M. Warnecke for invaluable comments and to Mr.W. Blair and Mr. D. J. Viery for critically re-viewing the manuscript. I would like to thankanonymous reviewers for their valuable sugges-tions on improving the paper.ReferencesChippindall, L. K. A. 1955. Miscanthidium.Meredith, D.（ed.）. The grasses and pasturesof South Africa, pp. 477―480. Central NewsAgency, Johannesburg.Clayton, W. D. 2002. Miscanthus Andersson.Pope, G. V. and Martins, E. S.（eds.）. FloraZambesiaca vol. 10 part 4, pp.8―10. RoyalBotanic Gardens, Kew.De Wet, J. M. J. 1960. Chromosome numbersand some morphological attributes of varioussouth African grasses. Amer. J. Bot. 47 : 44―49.Gibbs Russell, G. E. 1991. Miscanthus. Leistner,O. A.（ed.）. Grasses of southern Africa, p.221.National Botanic Gardens, Pretoria.Gibbs Russell, G. E., Reid, C., van Rooy, J. andSmook, L. 1985. List of species of southern Af-rican plants, ed. 2, part 1. 152 pp. BotanicalResearch Institute, Pretoria, Pretoria.Hodkinson, T. R., Chase, M. W., Lledo, M. D.,Salamin, N. and Renvoize, S. A. 2002. Phylo-genetics of Miscanthus, Saccharum and re-lated genera（Saccharinae, Andropogoneae,Poaceae）based on DNA sequences from ITSnuclear ribosomal DNA and plastid trnL in-tron and trnL-F intergenic spacers. J. Pl. Res.115 : 381―392.Hodkinson, T. R., Renvoize, S. A. and Chase, M.W. 1997. Systematics of Miscanthus. Aspectsof Applied Biology 49 : 189―198.Hoshino, T. and Davidse, G. 1988. Chromosomenumbers of grasses（Poaceae）from southernAfrica. 1. Ann. Missouri Bot. Gard. 75 : 866―873.Launert, E. 1970. Miscanthidium. Merxmüller,H.（ed.）. Prodromus einer flora von Süd-westafrica vol. 160, pp. 130―131. J. Cramer,Germany.Pilger, R. 1907. Gramineae Africanae VI. Engl.Jahrb. Syst. 39 : 597―601.Stapf, O. 1906. Plantae novae Daweanae inUganda lectae. J. Linn. Soc., Bot. 37 : 495―544.December 2004 J. Phytogeogr. Taxon. Vol. 52. No. 2－ 177 －（Received July 27, 2004 ; accepted December 6,2004）茨木 靖：アフリカ産イネ科植物の新組合わせMiscanthidium junceum subsp. teretifolium新組み合わせとしてMiscanthidium junceumsubsp. teretifolium を提唱した。M . teretifolium（Stapf）Stapfは，近年M . junceum（Stapf）Stapfのシノニムとされることが多いが，両者は分布域や包穎上の毛の状態などの点で異なっている。（〒770―8070 徳島市八万町文化の森総合公園 徳島県立博物館）植物地理・分類研究 第 52巻第 2号 2004年 12月－ 178 －

Miscanthidium junceum subsp. teretifolium （Poaceae）; a new combination for an African grass

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Miscanthidium junceum subsp. teretifolium （Poaceae）; a new combination for an African grass

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