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A revision of the Lower Cretaceous foraminiferal genus Falsogaudryinella from northwest Europe and Romania, and its relationship to Uvigerinammina

Abstract

We emend the definition of the foraminiferal genus Falsogaudryinella Bartenstein, 1977 based on observations of the type species, F. tealbyensis from the Barremian Lower Tealby Clay of Lincolnshire, U.K. The genus was described by Loeblich & Tappan (1987) as having initial triserial coiling which reduces to biserial and finally uniserial. However, topotype specimens display high trochospiral coiling in the microsphaeric generation, with at least four chambers in the initial whorl. The genus, therefore, does not belong in the family Verneuilinidae, but must be transferred to the Prolixoplectidae. The wall is solid, non-canaliculate. The connections between chambers are in the form of tubes that extend from the basal part of the chamber lumina toward a terminal aperture. This tubular connection is partially separated from the main part of the chamber lumina by a septum. The presence of this tubular connection in F. tealbyensis is closely analogous to that observed in the type species of Uvigerinammina Majzon, 1943. The two genera, therefore, are separated mainly on the basis of cement type, with Falsogaudryinella possessing calcareous cement and Uvigerinammina organic cement. We illustrate five species of Falsogaudryinella from the Barremian of Lincolnshire, the U.K. sector of the Central North Sea, and from the Barremian and the Albian of Romania (F. neagui Bartenstein, 1981, F. praemoesiana n.sp. F. tealbyensis (Bartenstein, 1956), F. xenogena n.sp. and F. moesiana (Neagu, 1966)). Our investigations reveal that upper Hauterivian to Barremian specimens from the North Sea that have been previously regarded as F. moesiana (e.g. King et al., 1989) in fact belong in a new species, Falsogaudryinella praemoesiana n.sp. A second new species, Falsogaudryinella xenogena n.sp. is described from the Barremian of the Central North Sea. Evolution within the mid- Cretaceous Falsogaudryinella group appears to progress by reduction of the terminal uniserial part, since the coiling in the stratigraphically youngest form (F. moesiana) is predominantly triserial. Our interpretations of the phylogeny of the Cretaceous Falsogaudryinella and Uvigerinammina lineages are presented

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