<p>Divergence estimates were based on (A) genetic population clusters and (B) traditional, morphology-based species circumscriptions. Posterior probabilities at nodes indicate support from the *BEAST analyses and speciation probabilities. Inset tables include divergence times for specified nodes, including the highest posterior density interval (HPD), using fixed substitution rates in four different scenarios: dates estimated using the ITS, nrLSU, RPB1 loci, independently; and the forth using rates from both the ITS nrLSU loci (Units: substitution/site/10<sup>9</sup> years). </p
