Aldrich, J.M.

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'''01. XXX] ENTO~rOLO(aCAL NE\VS ....~137Leiomyza in North America (Dipt., Drosophilidae.)By J. 1!. ALDRIClI, lJ. S. I)cparttl1cnt of .:\gricnlture, Bureauof EntoIllology, \ \~ashington, D. C.IIISTORICAL.Lcionl.\':::a ,,'as proposed h~T :\facqu:1rt in Tg35 for l\Ieigen's4'lgronly~a glabriclfla and lll('''z'i.aofa, \\"hie11. as l\lacquart oh-~tr\·cd. cliffcrcd too IllllCh on the ah~cllcc of vibrissae and ofztnterior frontal bristles to rCll1ain ill 4'} :;r()J/lyza; he also 111Cll-E~TO~[OLOGICAt XE\\"S. [:\Iay, '19tions approxinlated crossveins and the for\yard cun"ature ofthe fourth vein as generic characters, but the fonner of theseproved to be a Jl1istake.l\Ieigen in 1838 accepted the genus and added a third species,IJctcrollclfra scatopha!7ill<l F'allcn. \yhich he had hilllself pre-viously referred to ..~1!lr0111-,,=a. lIe fignres glabricllla. but thevenation is not quite alike in the t\\"o \\"ings, and he sho\ys afull cOTnplcnlcnt of ba~al cells \\"hich after\\'ard proYed anerror. He tllcntions as generic characters only the absenceof bristles on the anterior part of the front and the course ofthe fourth vein.\Vest\vood in 1840 placed 1.cio111-,,~a as a subgenus of ...:lgro-'11l.\'=a) defining it in a fc\\" \\"ords and repeating ).Iacquart'serror about the approxil11ation of the crossyeins ; he l11cntionedscatoplzaginQ as "typical species." although he probably had nothought of giving it any taxonoll1ic prolllinence ahoye theother t\\·o by so doing. As it \vas 110t originally included, itcould not be the type species in a t1lodern sense.Zctterstedt in I R48 gaye the best description of the genusyet published, evidently dra\ving the characters f rOl11 Fallen'stype of scatoplzar;iIlQ. \\"hich he also r('described; he also des-ignated this species as "T,::pus gCllcris. JJSchiner in 1864 \vas unable to give a satisfactory accountof the genus, his only n1aterial being a deterll1ined specinlenf ronl the I-Ialiday collection and one f rOlll \ \Tinncrtz, \\·hich\vere obviously not congeneric, yet both reselllblcci the (lesCrtp-tions. \Ve can no,,· decide that \\!innertz ,,"as right, hut~chiner could only adapt the earlier descriptions a fter afashion, leaving the genus 1110re confused than before.Rondani never attcll1ptcd to place the genus.Becker cleared up the Blatter very Bluch in 1902 by publish-ing the characters of the specinleB standing as type of laevi-p:ata in the l\Ieigen co11<.'ction. \\"ith a figure of the venation.This specinlen has the characters assigned by :\Icigen to thegenus, hut ~pccifically agrees better ,,·jth qlabriclfla, as it hasyelIo\v haltcres and the hind felllora not black at tip. Theylabrirlila type, or supposed type, is preserYcd \\"jth it but isno\v headless. I t has the satne \ving characters and is un-ENTO\IOLnCTC.\L XE\VS. 139doubtedly congeneric. T~('(ker Blakes the genus out as a per-fectly good 011C, difTering" frot11 ..~lstcia chietly in having a hindcrossvein and a longer (nor1l1al) second vcin~ and he placesit in Drosophilidae 11ext to that genns. 1;rol11 the Illaterial(~orth ~-\l11ericatl) t hat I ha \'e sectl. I enti rcly agree \vith this;\ve have a genus ~titl lllore nearly related. Sioa!ocssa, \vhichpossesses a hind CToss\'eil1, hence differs hardly at :111 except inits greatly ~h()rtel1ed ~ecot1d vein.Czerny in J()oJ gave S0111e notes on a ~pccinlent of scafoplza-.flina in Zetterstedt's collectioll, i11cluding- additional genericcharacters. 'This species is strict ly congeneric \\'1t11 the otber t\\'o,if not in fact an earlier nanle for .r;!abriclf!a. ~reigen in theoriginal description of locz"i.rJafa suggested that it lllig-ht be0nl)' a variant (:\h~inderung) of .rJlabriclfla. i\S no valid des-ignation of a genotype has heen tllade, glabriclfla is herebydesig-nated.\V'illiston in I ~l) (a) reported the genus f rOl11 North Alller-iea and placed it in his table of ./\grolllyzidae (I R<)6b) ; but in190R he oll1itted it, evidently not satisfied \\'ith his identification.:\Ielander in 1913, in his cxhausti\Te trcattllent of j\gro-111)'zidae and related fal11ilies~ tllentions Lcio111)'::"O as a genusunkno\\'n to hitn.I-iorenz Oldenherg in T9T 4 gives generic characters f raInIH1111erous speci Jl1ens and f ronl l~ecker and Czerny. I-Ie fi nelsthe flies in the forest in C;ertnany on exuding sap of trees, ontree f tlllgi, and on boards in the sun. II is SpeCit11enS sho\vconsiderable yariation in the amount of in fuscation of thefront, antennae and fel11ora. but seenl to agree \vith lac7'igola.\\rith this historical revie\\', I proceed to a description ofthe genus and t\\'o ne\\' North :\ tllerican species.GEXERIC CJI ..\R.\CTERS.Head nearly helllisphericaI. concave behind, the eye verylarge, bucca and para facial cxtrelllcly narro\v. ..Antennae or-dinary, inserted at about the llliddlc of the hr:ad in profile,arista bare or slightly pubescent. Face short, not carinate;a tllinute pair 0 f yihrissae prescnt. Front and face of equal\\·idth, less than one-third that of head; front flat, Slllooth, or-E~TO)[OLOGICAC NE\VS. [~Iay, '19hits tllOrc or less difTerentiated above for a short distance,and shortly above antennae \\~ith a transverse depression. T\vopairs of verticals, one of frontals, \\"hich are at the level ofthe median ocellus. l~ost-verticals absent (in ours, said to bedivergent by Oldenberg).Thorax and abdolllen in all the species black and highlypolished. C'haetotaxy of thorax: dc I. npl I, stpl I, SC I (and avery nlinute lateral pair, apparently absent in the fenlale ofsloSSOJlQ.c)" acrostichals in the :\orth ..-\lllerican species a verydelicate single ro\v of hairs, laterad of \yhich the surface isentirely bare to another ro\\' in the dc line. \\7"ing as figured,~econd basal and anal cells \\'anting, fourth vein gently curvedfor\vard from near the crossvein, so that the first posteriorcell is about t\Yo-thirds as \\,ide at apex as near its base.7~able of .\Tortll 6,111zcr;can Species.Front and hind felnora distinctly stouter than middle onessl~ssonae tl. SpaFront and hind fenlora of about the satne diallleter as the tniddleonesmelanderi n. sp.Leiomyza slossonae n. !'p.Front bro\vn, including an extension to the vertex on each side ofthe ocelli; ocellar triangle and the upper third of frontal orhits shiningblack; lower ('dge of front yellow; antennae yello\v. third joint, oval,slightly in f usca teo or not on upper edge; arista microscopicallypubescent, appearing hare under hand lens. Face. parafacials andbucca almost \vhite, sometirlles nlore yellowish, with a narro\v brownline usually perceptible separating the first t\\'o and bordering the sidesof the mouth cavity. Palpi yello\\': proboscis snlall, yello\v. I-Ialteresyellow. Legs including coxae \vholly yello\\' except last tarsal joint,which is bro\vn; front and hind fetnora thickened in both sexes, andtheir coxae and trochanters a little elongated. Length 1.8 mIn.TJ'pes, one 111ale (holotype), one felllale, l110unted on saInepin, fronl Franconia, 1\e\v Hatllpshire, collected by l\Trs. Slog..son and in her possession. Parat.vpes, one each frol11 Pulltnan,L.yndon and ..A.lnlota., \V"ashington, and one fronl 11oscow!\fountain, Idaho, all collected by Professor ~relander. Para--type fronl \Vashingtoll deposited in the United States National1\Illseum.\'01. XXX 1 ENTO\IOLO(;ICAI. NE\VS.Leiomyza melanderi n. sp. (Text Fig. 1, \ving).Slualler than slOSSOIlClC (1.5 111111.), all the fClllora slender; legs paleyellow. last tarsal j oint altn()~t black. Face not usually so whitish,Illore yellow. The specilllen f rOlll ~lontreal has the head darker, frontblackish-hrown, \\·ith a faint Inedian paler streak, lo\ver edge paleText Fig. L-Leiolll.n:a mdandr:1 ,. n. sp. Ri~ht winR. ulider side. X 46.yclIow, face and bucca quite dark yellow, third antennal joint abouthalf infuscated. Thi~ I can only regard as a dark fortn of the saInespecies, although the front looks di fferen t.Types, one l11ale, one fetnale, :\losco\v ~Iountain, Idaho;paraf),pcs, one each frot11 l\Iosco\v l\lountain, Idaho; Pull-tllan. (JIga, Everett and :-\ltllota, \ Vashillgton ; all the precedingcollected by Professor ~rclander, in \vhose possession the typesrenlain. ~A.lso one fet11ale paratype f rOlll l\lontreal, Canada,collected by Rev. Jas. Ouellet. 1\"0 paratypes from \Vashing-ton in the United States X ational :\1useunl.LITERATURE CITED.BESKI:R, 1902, Zeitsch. f. I-JYln. u. I)ipt., ii, 340.CZER~Y, 1903, \Vien. Ent. Zeit., xxii, 127.~IAcQcART, 1835, l-list. Kat. Dipt., ii, 605.~IEIGE~, 1838, Syst. Beschr. eur. Zweifl. Ins., vi, 394.~rEL.\~J)Er{, 1913, Jour. N. ·Y. Ent. Soc., xx, 228.OLDEXBERG, J914, .Arch. f. naturgesch., 8d ]ahrg., Ahth. A. Heft 2.P·36.S':HI~ER.. I86-t. Fauna Anstr., ii, 30<).\YEST\\"OOD, 1840 • Introd.~rod. Class., ii, app., 152.\\'ILLTSTOX, I896a, Ent. X ews, vii, 3~}~.(n., T8if)h. ~Iatlua] of X. :\. I)ipt., sec. ed., 104.In., 1<)08. ~[atll1at. 3d cd.ZETTERSTEDTJ 1R-tR, I)ipt. Scand., Yii, 2oiO.

Leiomyza in North America (Dipt., Drosophilidae.).

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Leiomyza in North America (Dipt., Drosophilidae.).

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