Two hundred and one species of Staphylinidae known to be found under bark or at sap of trees are listed. The possibility of these insects being important elements in the population dynamics of forest pests is discussed. An indication is given of the distribution and habitat of each species

Legner, E. F

Moore, Ian

English

ValpoScholar

The Great Lakes Entomologist 
Volume 10 
Number 4 - Winter 1977 Number 4 - Winter 
1977 
Article 2 
December 1977 
Staphylinidae from Under Bark and at Sap of Trees, a Preliminary 
Survey of Species Possibly Beneficial to Forestry (Coleoptera) 
E. F. Legner 
University of California 
Ian Moore 
University of California 
Follow this and additional works at: https://scholar.valpo.edu/tgle 
 Part of the Entomology Commons 
Recommended Citation 
Legner, E. F. and Moore, Ian 1977. "Staphylinidae from Under Bark and at Sap of Trees, a Preliminary 
Survey of Species Possibly Beneficial to Forestry (Coleoptera)," The Great Lakes Entomologist, vol 10 (4) 
Available at: https://scholar.valpo.edu/tgle/vol10/iss4/2 
This Peer-Review Article is brought to you for free and open access by the Department of Biology at ValpoScholar. 
It has been accepted for inclusion in The Great Lakes Entomologist by an authorized administrator of ValpoScholar. 
For more information, please contact a ValpoScholar staff member at scholar@valpo.edu. 
1977 THE GREAT LAKES ENTOMOLOGIST 
STAPHYLINIDAE FROM UNDER BARK AND AT SAP OF TREES, 
A PREI-IMINARY SURVEY OF SPECIES POSSIBLY 
BENEFICIAL TO FORESTRY (COLEOPTERA) 
E. F. Legner and Ian Moore 
ABSTRACT 
Ta-o hundred and one species of Staphylinidae known to be found under bark or at sap 
of trees are listed. The possibility of these insects being important elements in the popula- 
tion dynamics of forest pests is discussed. An indication is given of the distribution and 
habitat of each species. 
4 large percentage of the species of Staphylinidae whose ecology has been observed is 
predaceous Therefore, members of this family may play an important role in the popula- 
tion dynamics of pest insects in their special habitats. Considering the large body of litera- 
ture on the Staphylinidae, a very small amount of data is available on habitats, food habits 
and biologies of the approximately 30,000 described species. Consequently, information 
about species to be considered as candidates for biological control of pests in restricted 
habitats is difficult to obtain. The following list is intended to be an aid in that direction. 
Among Staphylinidae which are adapted for a subcortical habitat, most are well suited to 
that habitat by their depressed fonn. Many species known only from museum specimens are 
suspected to be subcortical by the dorso-ventral flattening of the body but actual data are 
lacking. 
Two subfamilies, Piestinae and Lispininae, have a larger percentage of their species with a 
subcortical habitat than l3e others, particularly as reported from the temperate fauna They 
have a ~ r y  high percentage of tropical species whose habitats are unreported; but, judging 
from their form are probably mostly subcortical. Some genera of other subfamilies such as 
in the Tach> porinae add quite a few- species to our list but are known often to be associated 
uith fungur It is ~ b l e  that their presence under the bark of dead trees is due to the 
presence of f u n y s  in that situation. Many such problems need to be investigated before the 
best use of any of these in- as biological control agents can be ascertained. 
The follou* lisi is compiled kom the examination of a large percentage of the existing 
literature, a wry small part of u f i i h  contains ecological data  This fact is attested by the 
small rize of the appended bibliographic list from which it was extracted. Only species 
actually reported as king arsociated a i th  trees, so encountered, or examined with such data 
are induded. 
The capital letters m parentheses folloaing the specific names indicate their reported 
distribution: A = Australian-Pacific, E = Ethiopian, H = Hawaiian, N = Nearctic, 0 = 
Oriental, T = Seotropical. The loa-er case letters indicate habitat: b = under bark, s = at sap. 
LIST OF SPECIES 
Subfamily ALEOCHARINAE 
Almchara ruficornis Gravenhorst ( P ,  b) Anomognathus cuspidatus Erichson (N, P, b) 
Aleodorus bilobatus Say (N, b) Atheta aequata Erichson ( P ,  b) 
l~ iv i s ion  of Biological Control, University of California, Riverside 92521. 
1
Legner and Moore: Staphylinidae from Under Bark and at Sap of Trees, a Preliminary
Published by ValpoScholar, 1977
174 THE GREAT LAKES ENTOMOLOGIST Vol. 10, No. 4 
A. autumnalis Erichson (P, b )  
A. bassicornis Mulsant & Rey (N,  P, b )  
A. celata Erichson (P, s) 
A. coriaria Kraatz (P, s) 
A. euryoptera Stephens (N, P, s) 
A. fungicola Thomson (P, s) 
A. linearis Gravenhorst (P, b )  
A. nigricornis Thomson (P, b ,  s) 
A. picipes Thomson (P, b )  
A. pilicornis'Thomson (P, b )  
A. sericea Mulsant & Rey (P, s) 
A. subcorticalis Cameron (0, b )  
A. triagulum Kraatz (P, s) 
Coenonica championi Cameron (0) 
C. mangpuensis Cameron (0, b )  
C. subopaca Cameron (0, b )  
C. puncticollis Kraatz (P, E, 0, s) 
Cordalia vestita Boheman (0) 
Cyphea curtula Erichson (P, b )  
Dadobia immersa Erichson (P, b )  
Geostiba ruficollis Erichson (P, b )  
Hornalata brunneicollis Cameron (0, b )  
H. plana Gyllenhal (P, b )  
H. uniformis Cameron (0, b) 
Leptusa angustata Aubd (P, b )  
Mimacrotona singulata Cameron (P, 0 ,  b )  
M. uniformis Cameron (0, b )  
Neoleptusa brunnea Cameron (0, b )  
Ocyusa incrassata Kraatz (P, b )  
Oligota variegata Blackburn (H, b )  
Oxypoda vittata Markel (P, s) 
Phloeopora corticalis Gravenhorst (P, b )  
P. diluta Sharp (H)  
P. montana Cameron (0) 
P. sublaevis Casey (N ,  b )  
P. reptans Gravenhorst (P, b )  
P. teres Gravenhorst (P, b )  
P. testacea Mamerheim (P, b )  
Placusa denticulata Shapp (P, s) 
P. fratercula Cameron (0, b )  
P. infima Erichson (P, b )  
Placusa pumilio Gravenhorst (P, b ,  s) 
Pseudatheta elegans Cameron (0) 
Silusa rubiginosa Erichson (P, b ,  s) 
Sipalia analis Gy llenhal (P, s) 
S. fumida Erichson (P, b )  
Stichoglossa corticina Erichson (P, 0 ,  b )  
S. indica Cameron (0, b )  
S. prolixa Gravenhorst (P, b )  
Tach~?usida gracilis Erichson (P, b )  
Thamiaraea cinnamomea Gravenhorst (P, s) 
T. hospita Markel (P, s) 
Thectura cuspidata Erichson (P, b )  
Troposilusa armata Cameron (0) 
Subfamily HYPOCYPHTINAE 
Anacyptus testaceus LeConte (N,  T ,  b )  
Subfamily LISPININAE 
Ancaeus pallescens Blackburn (H, b )  L. morugae Blackwelder (T ,  b )  
Cephalonyxum rambouseki Bierig ( T ,  b )  L. nigrifrons Fauvel ( T ,  b )  
~ l ~ v i l i s ~ i n u s  californicus ~ e ~ o n t e  (N,  b) L. stiiola Erichson ( T ,  b )  
Glvotoma trinidadensis Blackwelder (T.  b )  L. tenellus Erichson (T.  b )  
 hii in odes explicandus Sharp (H, b) 
Lispinus attenuatus Erichson (T,  b )  
L. bistriatus Fauvel(0, b )  
L. boxi Blackwelder (T ,  b )  
L. catena Sharp (T ,  b )  
L. danforthi Blackwelder (T ,  b )  
L. impar Blackwelder (T ,  b )  
L. impressicollis Motschulsky (H, b )  
L. irregularis Blackwelder (T ,  b )  
L. foveolus Blackwelder ( T ,  b )  
. , ,  
Neolispinodes exiguus Erichson ( T ,  b )  
N. politus Sharp ( T ,  b )  
N. rulomus Blackwelder (T ,  b)  
Renardia nigrellus LeConte (N,  b )  
Tetrapleurus himalayicus Cameron (0, b )  
Z4oracophorus brevicristatus Horn ( N ,  T ,  b )  
T. corticinus Motschulsky (P, b )  
T. guadalupensis Cameron (T ,  b )  
T. simplex Wendler (T ,  b )  
Subfamily MEGALOPSIDIINAE 
Megalopinus caelatus Gravenhorst (N ,  b )  M. smithi Cameron (T ,  b )  
2
The Great Lakes Entomologist, Vol. 10, No. 4 [1977], Art. 2
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THE GREAT LAKES ENTOMOLOGIST 
Subfamily MICROPEPLINAE 
Micropeplus brunneus Maklin (N) 
Subfamily OLISTHARINAE 
Olisthaerus megacephalus Zetterstedt (N, P, b) 0. substriatus Gyllenhal (N, P, b) 
Subfamily OMALIINAE 
Acrulia infhzfa Gyllenhal (P, b) 
A. tumidula Mgklin (N, b) 
Anthobium fectum Paykull (P, s) 
Arpedium cribratum Fauvel (N, b) 
Coryphium angusticolle Stephens (P, b) 
Hapalaraea brevicorne Erichson (P, b) 
H. f7avipenis Cameron (0 )  
H. gracilicorne Fairmaire (P, b) 
H. ioptma Stephens (P, b) 
H. pygmaea Gyllenhal (P, b) 
H. vile Erichson (P, b) 
Phloeonomus concolor Kraatz (P, b) 
P. minimus Erichson (P, b) 
P. peneti Thomson (P, b) 
P. planum Paykull (P, b, s) 
P. punctipenne Thomson (P, b) 
P. pusillurn Gravenhorst (N, P, b) 
Pycnoglypta lurida Gyllenhal (N, P. b) 
Xylodromus monilicorne Gyllenhal (P, b) 
Subfamily OXYTELINAE 
Oxytelus bledioides Blackburn (H, b) 
Subfamily PAEDERINAE 
Homeotarsus badium Gravenhorst (N, b) Lathrobium armatum Say (N, s) 
Subfamily PHLOEOCHARINAE 
Phloeochans subtilirs'ma 
Mannerheim 
(P, b) 
Subfamily PIESTINAE 
Hypotelus insulans Bierig (T, b) nigonurus asiaticus Reiche (P, b) 
Piestus peniclllatus (Dalman) (T, b) T. dilaticollis Van Dyke (N, b) 
Siagonium arnericmrum Melsheimer (N, b) T. caelatus LeConte (N, b) 
S. humerale Germar p, b) T. crotchei LeConte (N, b) 
S. punctatum LeConte (N, b) T. melleyi Mulsant & Rey (P, b) 
S. q u a d r b m e  Kirby (P, b) T. sharpi Blackwelder (N, b) 
S. stacesmithi Hatch (N, b) 
Subfamily PINOPHILINAE 
Palaminus testaceus Erichson (N, b) Pinophilus speculifions Cameron (0 )  
Subfamily PROTEININAE 
Megarthrus arcuatus Hatch (N) Metopsia clypeatum Miiller (P, b) 
Subfamily QUEDIINAE 
Ctenandropus nicriceps Cameron (0 ,  b) Heterothops occidentalis Casey (N, b) 
3
Legner and Moore: Staphylinidae from Under Bark and at Sap of Trees, a Preliminary
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176 THE GREAT LAKES ENTOMOLOGIST Vol. 10, No. 4 
H. pusio LeConte (N, b) Q. perigrinus Gravenhorst (N, b) 
Quedius brevicurnis Thomson (P, b) Q. rainieri Hatch (N, b) 
Q. charactus Hatch ( N ,  b) Q. rusricus Smetana (N, b) 
Q. cruenrus Olivier (P, b) Q. simplex Hatch (N, b) 
Q. f u m t u s  Stephens (P, b) Q. scitus Gravenhorst (N, b) 
Q. laevigatus Gyllenhal (N, P, b) Q. velox Smetana (N, b) 
Q. longicornis Kraatz (P, b) Q. xanihopus Erichson (P, b) 
Subfamily STAPHYLININAE 
Belonuchus ceylonicus Cameron ( 0 ,  b) H. laevis Blackwelder (T, b) 
B. formosus Gravenhorst (N, T ,  s) Laetzrlonthus laetulus Say (N, b) 
B. gardeneri Cameron ( 0 ,  b) Paracraspedomerus speculifrons Fauvel (A, b) 
B. rufoniger Fauvel(0,  b) Philonrhus chalceus Stephens (N, P, s) 
Diplosrictus chenui Perroud (A, b) ,P. crestonensis Hatch (N, b) 
Erichsoniusfrarerculus Horn (N, b) P. flavocinctus Motschulsky (A, b) 
Holisus guildingii Erichson (T, b) P. imitator Cameron ( 0 ,  b) 
Subfamily TACHYPORINAE 
Coproporus ebonus Blackwelder (T, b) 
C. hepaticus Erichson (T, b) 
C. lecontei Blackwelder (N, b) 
C. monticola Cameron ( 0 ,  b) 
C. ventriculus Say (N, b) 
Lordithon intrusus Horn (N, b) 
Sepedophilus basalis Erichson (N, b) 
S. beckeri Campbell (N, b) 
S. castaneus Horn (N, b) 
S. cincrus Erichson (N, b) 
S. ctenidialis Campbell (N, b) 
S. hermni  Campbell (N, b) 
S. kiteleyi Campbell (N, b) 
S. littoreus Linnd (N, P, b) 
S. marshami Stephens (N, P, b) 
S. ocultus Casey (N, b) 
S. opicus Say (N, b) 
S. scriptus Horn (N, b) 
S. srnetami Campbell (N, b) 
S. stephani Campbell (N, b) 
S. testaceus Fabricius (N, P, b) 
Tachinus angustatus Horn (N, b) 
T. debilis Horn (N, b) 
Subfamily TRICHOPHYINAE 
Trichophya pilicornis Gyllenhal (N, P, 0 ,  b) 
Subfamily XANTHOLININAE 
Atrecus affinis Paykull (P ,  b) Nudobius cephalus Say (N, b) 
A. alternuns Gravenhorst (P, b) N, collaris Erichson (P, b) 
A. longiceps Fauvel (P, b) N. lentus Erichson (P, b) 
A. pilicornis Paykull (P, b) Oligolinus filarius Erichson (T, b) 
Hyponygrus illucens Erichson (T, b) 0. testaceipennis Fauvel (P, b) 
H. glaber Nordmann (P, b) Srenolinus macrothricus Bierig (T, b) 
H. gularis LeConte (N, b) Zeteotomus brevicornis Erichson (P, b) 
Microlinus pusio LeConte (N, T, B) 
Subfamily XANTHOPYGINAE 
Holisomimus parvus Cameron ( 0 ,  b) 
4
The Great Lakes Entomologist, Vol. 10, No. 4 [1977], Art. 2
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THE GREAT LAKES ENTOMOLOGIST 
REFERENCES 
Blackwelder, R. E. 194 1. A monograph of the genus Trigonurus (Coleoptera: Staphylini- 
dae). Amer. Mus. Nov. 1124: 1-13. 
1943. Monograph of the West Indian beetles of the family Staphylinidae. 
Bull. U.S. Nat. Mus. 185:l-188. 
Blatchley, W. S. 1910. The Coleoptera or beetles (exclusive of the Rhycophora) known to  
occw in Indiana with bibliography and description of new species. Ind. Geol. & Nat. Res., 
Bull. 1: 1-1386. 
Cameron, M. 1930. The fauna of British India, including Ceylon and Burma Coleoptera, 
Staphylinidae, vol. 1. London. 
-- 1931. The fauna of British India, including Ceylon and Burma. Coleoptera, 
Staphylinidae, Vol. 2. London. 
1932. The fauna of British India, including Ceylon and Burma. Coleoptera, 
Staphylinidae, vol. 3. London. 
1939. The fauna of British India, including Ceylon and Burma. Coleoptera, 
Staphylinidae, vol. 4, pt. 1. London. 
1939. The fauna of British India, including Ceylon and Burma. Coleoptera, 
- ~ 
Staphylinjdae, vol. 4, pt. 2. London. 
Campbell. J. M. 1968. New World Micropeplinae. Can. Entomol. 100:225-267. 
- .  
- 1973. A revision of the GnLs Tachinus (Coleoptera: Staphylinidae) of North 
and Central America. Mem. Entomol. Soc. Can. 90:l-137. 
1975. A revision of the genera Coproporus and Cilea (Coleoptera: 
Staphylinidae) of America north of Mexico. Can. Entomol. 107:175-216. 
1976. A revision of the genus Sepedophilus Gistel (Co1eoptera:Staphylinidae) 
of America north of Mexico. Mem. Entomol. Soc. Can. 99: 1-89. 
Coiffait, H. 1972. Col6optires Staphylinidae de la Palkarctic occidentale. I. Generalit&. 
Sous-families: Xantholininae et Leptotyphlinae. Publ. Nouv. Rev. d'Entomol. 2: 1-651. 
Fagel, G. 1968. Contribution a la connaisances des Staphylinidae. CVI. Remarques diverus 
sw les especes de la rCgion paliarctique occidentale. Bull. Soc. Roy. Entomol. Belg. 
104: 189-204. 
Fowler, \V. W. 1888. The Coleoptera of the British Islands. A descriptive account of the 
families, genera and species indigenous to Great Britain and Ireland, with notes as to 
localities, habitats, etc., vol. 2, Staphylinidae. London. 
Ganglbauer, L. 1885. Die Kifer von Mitteleuropa. Die Kgfer der ~sterreichish-ungarischen 
Monarchie Deutschlands, der Schweiz, sowie des Franzosischen und Italienischen 
Alpengebietes, vol. 2, Familienreihe Staphylinoidea, 1. Theil: Staphylinidae, Pselaphidae. 
Vienna. 
Hatch, M. 1957. The beetles of the Pacific Northwest. Part 11: Staphyliniformia. Univ. Wash. 
Press Publ. Biol. 16: 1-384. 
LeConte, J. L. 1877. On certain genera of Staphylinidae, Oxytelini, Piestidae and Micro- 
peplidae as represented in the fauna of the United States. Trans. Amer. Entomol. Soc. 
6:213-252. 
Moore, Ian. 1975. The distribution of Siagonium (Coleoptera: Staphylinidae) in North 
America J. N. Y. Entomol. Soc. 48:96-100. 
Sharp, D. 1880. On some Coleoptera from the Hawaiian Islands. Trans. Entomol. Soc. 
London 1880:37-54. 
1885. In: Blackburn and Sharp, Memoirs on the Coleoptera of the Hawaiian 
Islands. Sci. Tran. Royal Dublin Soc., Ser. 2, 3: 119-300. 
Sha~p,  D. and H. Scott. 1908. Fauna Hawaiiensis. Coleoptera (various) 3:367-578. 
Scheerpeltz, 0. 1966. Ergebnisse der ~sterreichischen Neukaledonia-Expedition 1965- 
Staphyliniden. Ann. Natur. Mus. Wien 69:393418. 
Smetana, A. 1971. Revision of the tribe Quediini of America north of Mexico (Coleoptera: 
Staphylinidae). Mem. Entomol. Soc. Can. 79: 1-303. 
1976. Revision of the tribe Quediini of America north of Mexico (Coleoptera: 
Staphylinidae). Supplementum 3. Can. Entomol. 108: 169-184. 
Struble, G. E. 1930. The biology of certain Coleoptera associated with bark beetles in 
western yellow pine. Univ. Calif. Publ. Entomol. 5:107-134. 
5
Legner and Moore: Staphylinidae from Under Bark and at Sap of Trees, a Preliminary
Published by ValpoScholar, 1977


Staphylinidae from Under Bark and at Sap of Trees, a Preliminary Survey of Species Possibly Beneficial to Forestry (Coleoptera)

Valparaiso University

The Great Lakes Entomologist Volume 10 Number 4 - Winter 1977 Number 4 - Winter 1977 Article 2 December 1977 Staphylinidae from Under Bark and at Sap of Trees, a Preliminary Survey of Species Possibly Beneficial to Forestry (Coleoptera) E. F. Legner University of California Ian Moore University of California Follow this and additional works at: https://scholar.valpo.edu/tgle  Part of the Entomology Commons Recommended Citation Legner, E. F. and Moore, Ian 1977. "Staphylinidae from Under Bark and at Sap of Trees, a Preliminary Survey of Species Possibly Beneficial to Forestry (Coleoptera)," The Great Lakes Entomologist, vol 10 (4) Available at: https://scholar.valpo.edu/tgle/vol10/iss4/2 This Peer-Review Article is brought to you for free and open access by the Department of Biology at ValpoScholar. It has been accepted for inclusion in The Great Lakes Entomologist by an authorized administrator of ValpoScholar. For more information, please contact a ValpoScholar staff member at scholar@valpo.edu. 1977 THE GREAT LAKES ENTOMOLOGIST STAPHYLINIDAE FROM UNDER BARK AND AT SAP OF TREES, A PREI-IMINARY SURVEY OF SPECIES POSSIBLY BENEFICIAL TO FORESTRY (COLEOPTERA) E. F. Legner and Ian Moore ABSTRACT Ta-o hundred and one species of Staphylinidae known to be found under bark or at sap of trees are listed. The possibility of these insects being important elements in the popula- tion dynamics of forest pests is discussed. An indication is given of the distribution and habitat of each species. 4 large percentage of the species of Staphylinidae whose ecology has been observed is predaceous Therefore, members of this family may play an important role in the popula- tion dynamics of pest insects in their special habitats. Considering the large body of litera- ture on the Staphylinidae, a very small amount of data is available on habitats, food habits and biologies of the approximately 30,000 described species. Consequently, information about species to be considered as candidates for biological control of pests in restricted habitats is difficult to obtain. The following list is intended to be an aid in that direction. Among Staphylinidae which are adapted for a subcortical habitat, most are well suited to that habitat by their depressed fonn. Many species known only from museum specimens are suspected to be subcortical by the dorso-ventral flattening of the body but actual data are lacking. Two subfamilies, Piestinae and Lispininae, have a larger percentage of their species with a subcortical habitat than l3e others, particularly as reported from the temperate fauna They have a ~ r y  high percentage of tropical species whose habitats are unreported; but, judging from their form are probably mostly subcortical. Some genera of other subfamilies such as in the Tach> porinae add quite a few- species to our list but are known often to be associated uith fungur It is ~ b l e  that their presence under the bark of dead trees is due to the presence of f u n y s  in that situation. Many such problems need to be investigated before the best use of any of these in- as biological control agents can be ascertained. The follou* lisi is compiled kom the examination of a large percentage of the existing literature, a wry small part of u f i i h  contains ecological data  This fact is attested by the small rize of the appended bibliographic list from which it was extracted. Only species actually reported as king arsociated a i th  trees, so encountered, or examined with such data are induded. The capital letters m parentheses folloaing the specific names indicate their reported distribution: A = Australian-Pacific, E = Ethiopian, H = Hawaiian, N = Nearctic, 0 = Oriental, T = Seotropical. The loa-er case letters indicate habitat: b = under bark, s = at sap. LIST OF SPECIES Subfamily ALEOCHARINAE Almchara ruficornis Gravenhorst ( P ,  b) Anomognathus cuspidatus Erichson (N, P, b) Aleodorus bilobatus Say (N, b) Atheta aequata Erichson ( P ,  b) l~ iv i s ion  of Biological Control, University of California, Riverside 92521. 1Legner and Moore: Staphylinidae from Under Bark and at Sap of Trees, a PreliminaryPublished by ValpoScholar, 1977174 THE GREAT LAKES ENTOMOLOGIST Vol. 10, No. 4 A. autumnalis Erichson (P, b )  A. bassicornis Mulsant & Rey (N,  P, b )  A. celata Erichson (P, s) A. coriaria Kraatz (P, s) A. euryoptera Stephens (N, P, s) A. fungicola Thomson (P, s) A. linearis Gravenhorst (P, b )  A. nigricornis Thomson (P, b ,  s) A. picipes Thomson (P, b )  A. pilicornis'Thomson (P, b )  A. sericea Mulsant & Rey (P, s) A. subcorticalis Cameron (0, b )  A. triagulum Kraatz (P, s) Coenonica championi Cameron (0) C. mangpuensis Cameron (0, b )  C. subopaca Cameron (0, b )  C. puncticollis Kraatz (P, E, 0, s) Cordalia vestita Boheman (0) Cyphea curtula Erichson (P, b )  Dadobia immersa Erichson (P, b )  Geostiba ruficollis Erichson (P, b )  Hornalata brunneicollis Cameron (0, b )  H. plana Gyllenhal (P, b )  H. uniformis Cameron (0, b) Leptusa angustata Aubd (P, b )  Mimacrotona singulata Cameron (P, 0 ,  b )  M. uniformis Cameron (0, b )  Neoleptusa brunnea Cameron (0, b )  Ocyusa incrassata Kraatz (P, b )  Oligota variegata Blackburn (H, b )  Oxypoda vittata Markel (P, s) Phloeopora corticalis Gravenhorst (P, b )  P. diluta Sharp (H)  P. montana Cameron (0) P. sublaevis Casey (N ,  b )  P. reptans Gravenhorst (P, b )  P. teres Gravenhorst (P, b )  P. testacea Mamerheim (P, b )  Placusa denticulata Shapp (P, s) P. fratercula Cameron (0, b )  P. infima Erichson (P, b )  Placusa pumilio Gravenhorst (P, b ,  s) Pseudatheta elegans Cameron (0) Silusa rubiginosa Erichson (P, b ,  s) Sipalia analis Gy llenhal (P, s) S. fumida Erichson (P, b )  Stichoglossa corticina Erichson (P, 0 ,  b )  S. indica Cameron (0, b )  S. prolixa Gravenhorst (P, b )  Tach~?usida gracilis Erichson (P, b )  Thamiaraea cinnamomea Gravenhorst (P, s) T. hospita Markel (P, s) Thectura cuspidata Erichson (P, b )  Troposilusa armata Cameron (0) Subfamily HYPOCYPHTINAE Anacyptus testaceus LeConte (N,  T ,  b )  Subfamily LISPININAE Ancaeus pallescens Blackburn (H, b )  L. morugae Blackwelder (T ,  b )  Cephalonyxum rambouseki Bierig ( T ,  b )  L. nigrifrons Fauvel ( T ,  b )  ~ l ~ v i l i s ~ i n u s  californicus ~ e ~ o n t e  (N,  b) L. stiiola Erichson ( T ,  b )  Glvotoma trinidadensis Blackwelder (T.  b )  L. tenellus Erichson (T.  b )   hii in odes explicandus Sharp (H, b) Lispinus attenuatus Erichson (T,  b )  L. bistriatus Fauvel(0, b )  L. boxi Blackwelder (T ,  b )  L. catena Sharp (T ,  b )  L. danforthi Blackwelder (T ,  b )  L. impar Blackwelder (T ,  b )  L. impressicollis Motschulsky (H, b )  L. irregularis Blackwelder (T ,  b )  L. foveolus Blackwelder ( T ,  b )  . , ,  Neolispinodes exiguus Erichson ( T ,  b )  N. politus Sharp ( T ,  b )  N. rulomus Blackwelder (T ,  b)  Renardia nigrellus LeConte (N,  b )  Tetrapleurus himalayicus Cameron (0, b )  Z4oracophorus brevicristatus Horn ( N ,  T ,  b )  T. corticinus Motschulsky (P, b )  T. guadalupensis Cameron (T ,  b )  T. simplex Wendler (T ,  b )  Subfamily MEGALOPSIDIINAE Megalopinus caelatus Gravenhorst (N ,  b )  M. smithi Cameron (T ,  b )  2The Great Lakes Entomologist, Vol. 10, No. 4 [1977], Art. 2https://scholar.valpo.edu/tgle/vol10/iss4/2THE GREAT LAKES ENTOMOLOGIST Subfamily MICROPEPLINAE Micropeplus brunneus Maklin (N) Subfamily OLISTHARINAE Olisthaerus megacephalus Zetterstedt (N, P, b) 0. substriatus Gyllenhal (N, P, b) Subfamily OMALIINAE Acrulia infhzfa Gyllenhal (P, b) A. tumidula Mgklin (N, b) Anthobium fectum Paykull (P, s) Arpedium cribratum Fauvel (N, b) Coryphium angusticolle Stephens (P, b) Hapalaraea brevicorne Erichson (P, b) H. f7avipenis Cameron (0 )  H. gracilicorne Fairmaire (P, b) H. ioptma Stephens (P, b) H. pygmaea Gyllenhal (P, b) H. vile Erichson (P, b) Phloeonomus concolor Kraatz (P, b) P. minimus Erichson (P, b) P. peneti Thomson (P, b) P. planum Paykull (P, b, s) P. punctipenne Thomson (P, b) P. pusillurn Gravenhorst (N, P, b) Pycnoglypta lurida Gyllenhal (N, P. b) Xylodromus monilicorne Gyllenhal (P, b) Subfamily OXYTELINAE Oxytelus bledioides Blackburn (H, b) Subfamily PAEDERINAE Homeotarsus badium Gravenhorst (N, b) Lathrobium armatum Say (N, s) Subfamily PHLOEOCHARINAE Phloeochans subtilirs'ma Mannerheim (P, b) Subfamily PIESTINAE Hypotelus insulans Bierig (T, b) nigonurus asiaticus Reiche (P, b) Piestus peniclllatus (Dalman) (T, b) T. dilaticollis Van Dyke (N, b) Siagonium arnericmrum Melsheimer (N, b) T. caelatus LeConte (N, b) S. humerale Germar p, b) T. crotchei LeConte (N, b) S. punctatum LeConte (N, b) T. melleyi Mulsant & Rey (P, b) S. q u a d r b m e  Kirby (P, b) T. sharpi Blackwelder (N, b) S. stacesmithi Hatch (N, b) Subfamily PINOPHILINAE Palaminus testaceus Erichson (N, b) Pinophilus speculifions Cameron (0 )  Subfamily PROTEININAE Megarthrus arcuatus Hatch (N) Metopsia clypeatum Miiller (P, b) Subfamily QUEDIINAE Ctenandropus nicriceps Cameron (0 ,  b) Heterothops occidentalis Casey (N, b) 3Legner and Moore: Staphylinidae from Under Bark and at Sap of Trees, a PreliminaryPublished by ValpoScholar, 1977176 THE GREAT LAKES ENTOMOLOGIST Vol. 10, No. 4 H. pusio LeConte (N, b) Q. perigrinus Gravenhorst (N, b) Quedius brevicurnis Thomson (P, b) Q. rainieri Hatch (N, b) Q. charactus Hatch ( N ,  b) Q. rusricus Smetana (N, b) Q. cruenrus Olivier (P, b) Q. simplex Hatch (N, b) Q. f u m t u s  Stephens (P, b) Q. scitus Gravenhorst (N, b) Q. laevigatus Gyllenhal (N, P, b) Q. velox Smetana (N, b) Q. longicornis Kraatz (P, b) Q. xanihopus Erichson (P, b) Subfamily STAPHYLININAE Belonuchus ceylonicus Cameron ( 0 ,  b) H. laevis Blackwelder (T, b) B. formosus Gravenhorst (N, T ,  s) Laetzrlonthus laetulus Say (N, b) B. gardeneri Cameron ( 0 ,  b) Paracraspedomerus speculifrons Fauvel (A, b) B. rufoniger Fauvel(0,  b) Philonrhus chalceus Stephens (N, P, s) Diplosrictus chenui Perroud (A, b) ,P. crestonensis Hatch (N, b) Erichsoniusfrarerculus Horn (N, b) P. flavocinctus Motschulsky (A, b) Holisus guildingii Erichson (T, b) P. imitator Cameron ( 0 ,  b) Subfamily TACHYPORINAE Coproporus ebonus Blackwelder (T, b) C. hepaticus Erichson (T, b) C. lecontei Blackwelder (N, b) C. monticola Cameron ( 0 ,  b) C. ventriculus Say (N, b) Lordithon intrusus Horn (N, b) Sepedophilus basalis Erichson (N, b) S. beckeri Campbell (N, b) S. castaneus Horn (N, b) S. cincrus Erichson (N, b) S. ctenidialis Campbell (N, b) S. hermni  Campbell (N, b) S. kiteleyi Campbell (N, b) S. littoreus Linnd (N, P, b) S. marshami Stephens (N, P, b) S. ocultus Casey (N, b) S. opicus Say (N, b) S. scriptus Horn (N, b) S. srnetami Campbell (N, b) S. stephani Campbell (N, b) S. testaceus Fabricius (N, P, b) Tachinus angustatus Horn (N, b) T. debilis Horn (N, b) Subfamily TRICHOPHYINAE Trichophya pilicornis Gyllenhal (N, P, 0 ,  b) Subfamily XANTHOLININAE Atrecus affinis Paykull (P ,  b) Nudobius cephalus Say (N, b) A. alternuns Gravenhorst (P, b) N, collaris Erichson (P, b) A. longiceps Fauvel (P, b) N. lentus Erichson (P, b) A. pilicornis Paykull (P, b) Oligolinus filarius Erichson (T, b) Hyponygrus illucens Erichson (T, b) 0. testaceipennis Fauvel (P, b) H. glaber Nordmann (P, b) Srenolinus macrothricus Bierig (T, b) H. gularis LeConte (N, b) Zeteotomus brevicornis Erichson (P, b) Microlinus pusio LeConte (N, T, B) Subfamily XANTHOPYGINAE Holisomimus parvus Cameron ( 0 ,  b) 4The Great Lakes Entomologist, Vol. 10, No. 4 [1977], Art. 2https://scholar.valpo.edu/tgle/vol10/iss4/2THE GREAT LAKES ENTOMOLOGIST REFERENCES Blackwelder, R. E. 194 1. A monograph of the genus Trigonurus (Coleoptera: Staphylini- dae). Amer. Mus. Nov. 1124: 1-13. 1943. Monograph of the West Indian beetles of the family Staphylinidae. Bull. U.S. Nat. Mus. 185:l-188. Blatchley, W. S. 1910. The Coleoptera or beetles (exclusive of the Rhycophora) known to  occw in Indiana with bibliography and description of new species. Ind. Geol. & Nat. Res., Bull. 1: 1-1386. Cameron, M. 1930. The fauna of British India, including Ceylon and Burma Coleoptera, Staphylinidae, vol. 1. London. -- 1931. The fauna of British India, including Ceylon and Burma. Coleoptera, Staphylinidae, Vol. 2. London. 1932. The fauna of British India, including Ceylon and Burma. Coleoptera, Staphylinidae, vol. 3. London. 1939. The fauna of British India, including Ceylon and Burma. Coleoptera, Staphylinidae, vol. 4, pt. 1. London. 1939. The fauna of British India, including Ceylon and Burma. Coleoptera, - ~ Staphylinjdae, vol. 4, pt. 2. London. Campbell. J. M. 1968. New World Micropeplinae. Can. Entomol. 100:225-267. - .  - 1973. A revision of the GnLs Tachinus (Coleoptera: Staphylinidae) of North and Central America. Mem. Entomol. Soc. Can. 90:l-137. 1975. A revision of the genera Coproporus and Cilea (Coleoptera: Staphylinidae) of America north of Mexico. Can. Entomol. 107:175-216. 1976. A revision of the genus Sepedophilus Gistel (Co1eoptera:Staphylinidae) of America north of Mexico. Mem. Entomol. Soc. Can. 99: 1-89. Coiffait, H. 1972. Col6optires Staphylinidae de la Palkarctic occidentale. I. Generalit&. Sous-families: Xantholininae et Leptotyphlinae. Publ. Nouv. Rev. d'Entomol. 2: 1-651. Fagel, G. 1968. Contribution a la connaisances des Staphylinidae. CVI. Remarques diverus sw les especes de la rCgion paliarctique occidentale. Bull. Soc. Roy. Entomol. Belg. 104: 189-204. Fowler, \V. W. 1888. The Coleoptera of the British Islands. A descriptive account of the families, genera and species indigenous to Great Britain and Ireland, with notes as to localities, habitats, etc., vol. 2, Staphylinidae. London. Ganglbauer, L. 1885. Die Kifer von Mitteleuropa. Die Kgfer der ~sterreichish-ungarischen Monarchie Deutschlands, der Schweiz, sowie des Franzosischen und Italienischen Alpengebietes, vol. 2, Familienreihe Staphylinoidea, 1. Theil: Staphylinidae, Pselaphidae. Vienna. Hatch, M. 1957. The beetles of the Pacific Northwest. Part 11: Staphyliniformia. Univ. Wash. Press Publ. Biol. 16: 1-384. LeConte, J. L. 1877. On certain genera of Staphylinidae, Oxytelini, Piestidae and Micro- peplidae as represented in the fauna of the United States. Trans. Amer. Entomol. Soc. 6:213-252. Moore, Ian. 1975. The distribution of Siagonium (Coleoptera: Staphylinidae) in North America J. N. Y. Entomol. Soc. 48:96-100. Sharp, D. 1880. On some Coleoptera from the Hawaiian Islands. Trans. Entomol. Soc. London 1880:37-54. 1885. In: Blackburn and Sharp, Memoirs on the Coleoptera of the Hawaiian Islands. Sci. Tran. Royal Dublin Soc., Ser. 2, 3: 119-300. Sha~p,  D. and H. Scott. 1908. Fauna Hawaiiensis. Coleoptera (various) 3:367-578. Scheerpeltz, 0. 1966. Ergebnisse der ~sterreichischen Neukaledonia-Expedition 1965- Staphyliniden. Ann. Natur. Mus. Wien 69:393418. Smetana, A. 1971. Revision of the tribe Quediini of America north of Mexico (Coleoptera: Staphylinidae). Mem. Entomol. Soc. Can. 79: 1-303. 1976. Revision of the tribe Quediini of America north of Mexico (Coleoptera: Staphylinidae). Supplementum 3. Can. Entomol. 108: 169-184. Struble, G. E. 1930. The biology of certain Coleoptera associated with bark beetles in western yellow pine. Univ. Calif. Publ. Entomol. 5:107-134. 5Legner and Moore: Staphylinidae from Under Bark and at Sap of Trees, a PreliminaryPublished by ValpoScholar, 1977

E. F Legner

Ian Moore

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Staphylinidae from Under Bark and at Sap of Trees, a Preliminary Survey of Species Possibly Beneficial to Forestry (Coleoptera)

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