The family Tenebrionidae, the darkling beetles, containing more than 14,000 species, is found primarily in the tropics and arid regions. As would therefore be expected, Michigan\u27s cool, moist environment is not attractive to very many tenebrionids. This list contains 50 identified species and six previously recorded species that were not confirmed by me

Spilman, T.J.

English

ValpoScholar

The Great Lakes Entomologist 
Volume 6 
Number 3 -- Fall 1973 Number 3 -- Fall 1973 Article 2 
August 2017 
A List of the Tenebrionidae of Michigan (Coleoptera) 
T.J. Spilman 
US Department of Agriculture 
Follow this and additional works at: https://scholar.valpo.edu/tgle 
 Part of the Entomology Commons 
Recommended Citation 
Spilman, T.J. 2017. "A List of the Tenebrionidae of Michigan (Coleoptera)," The Great Lakes Entomologist, 
vol 6 (3) 
Available at: https://scholar.valpo.edu/tgle/vol6/iss3/2 
This Peer-Review Article is brought to you for free and open access by the Department of Biology at ValpoScholar. 
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1973 THE GREAT LAKES ENTOMOLOGIST 
A LIST OF THE TENEBRIONIDAE OF MICHIGAN (COLEOPTERA) 
T. J. Spilman 
Entomology Research Division, Agricultural Research Service, 
United States Departmeni of Agriculture, Washington, D . c . ~  
The family Tenebrionidae, the darkling beetles, containing more than 14,000 species, 
is found primarily in the tropics and arid regions. As would therefore be expected, 
Michigan's cool, moist environment is not attractive to very many tenebrionids. This list 
contains 50 identified species and six previously recorded species that were not confirmed 
by me. 
Previously 45 species of Tenebrionidae had been recorded in Michigan in various lists. 
I have not attempted to find all literature records, but I have seen the principal faunal 
lists of Coleoptera. The three areas of the state, Lower Peninsula, Upper Peninsula, and 
Isle Royale, are used here to summarize previous lists. In the Lower Peninsula, Hubbard 
and Schwarz (1878b) recorded 32 species, Townsend (1889) recorded three species, 
Andrews (1916) recorded 21 species from the Charity Islands (Arenac County) in the 
mouth of Saginaw Bay, and Hatch (1924) recorded 11 species from Charlevoix County. 
In the Upper Peninsula, Hubbard and Schwarz (1878a) recorded 16 species from the 
Lake Superior area, and Andrews (1923) recorded 13 species from Whitefish Point. On 
Isle Royale, Adams (1909) recorded one species. After duplications in these lists are 
accounted for, the Lower Peninsula had 34 species, the Upper Peninsula had 25, Isle 
Royale, of course, had only one, and the whole state had 45. Of the 45 species, six are 
not recorded in my list; those six are discussed in the Appendix. In my list of 50 
identified species, 46 occur in the Lower Peninsula, 26 occur in the Upper Peninsula, 
none is recorded from Isle Royale, and one species had only a state label on the 
specimen. 
This list is based on specimens I have examined from the following institutions: 
University of Michigan Museum of Zoology, Ann Arbor, lent by Richard D. Alexander; 
Michigan State University, East Lansing, lent by Roland L. Fischer and the late R. R. 
Dreisbach; U.S. National Museum of Natural History, Washington, D.C. In addition to 
!locality records from my identifications, I have included localities published by C. A. 
Triplehorn in his 1965 revision of the Diaperini. 
DISTRIBUTIONAL DATA 
The basic unit of distributional information is the county. Counties are numbered by 
tiers, beginning in the southwestern corner of the state. The counties in the Lower 
Peninsula are numbered from 10 to 113, those in the Upper Peninsula from 200 to  214, 
and Isle Royale 300. The system permits condensation of data and, after a little 
familiarity, easy visualization of county position in the state by its number alone (see 
Fig. 1). The numbering of the counties is as follows: 
10-Berrien 25-Wayne 44-Shiawassee 
11-Cass 30-Allegan 45-Genesee 
12-St. Joseph 31-Barry 46-Lapeer 
13-Branch 3 2-Eaton 47-St. Clair 
14-Hillsdale 33-Ingham 50-Muskegon 
15 -Lenawee 34-Livingston 5 1-Montcalm 
16-Monroe 35 -Oakland 52-Gratiot 
20-Van Buren 36-Macomb 53-Saginaw 
21-Kalamazoo 40-Ottawa 54-Tuscola 
22-Calhoun 41-Kent 55 -Sanilac 
23-Jackson 42-Ionia 60-Oceana 
24-Washtenaw 43-Clinton 6 1 -Newaygo 
l ~ a i l i n ~  address: Department of Entomology, Smithsonian Institution, Washington, 
D.C. 20560. 
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Spilman: A List of the Tenebrionidae of Michigan (Coleoptera)
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THE GREAT LAKES ENTOMOLOGIST Vol. 6, No. 3 
Fig. 1. The counties of the State of Michigan indicated by numbers. For explanation, see 
text page 85. 
75 -Arenac 
80-Manistee 
8 1 -Wexford 
82-Missaukee 
83-Roscommon 
84-Ogemaw 
85-IOSCO 
90-Benzie 
91 -Grand Traverse 
92-Kalkaska 
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THE GREAT LAKES ENTOMOLOGIST 
112-Cheboygan 
113 -Presque Isle 
200-Gogebic 
201 -0ntonagon 
202-Houghton 
203-Keweenaw 
Subfamily ASIDINAE 
Tribe ZOPHERINI 
Genus PHELLOPSIS LeConte 
obcordata (Kirby). 112 207 214. July 5 to September 16. Habitat: under bark of dead 
trees. 
Subfamily TENEBRIONINAE 
Tribe PEDININI 
Genus BLAPSTINUS Dejean 
The species are often found in debris on ground, also in grasses. Some species in other 
parts of the United States occasionally build up in great numbers and damage truck 
crops. 
metallicus (Fabricius). 10  25 31 35 47 66 83 85 93 95 100 110 213. April 24 to 
September 3. 
moestus Melsheimer. 10 25. July 24. 
Tribe BOLITOPHAGINI 
Genus BOLITOTHERUS Candkze 
cornutus (Panzer). 10 20 21 24 25 33 34 35 36 40 44 45 66 75 91 100 110 200 201 
204 207 210 21 1 213 214. April 5 to September 26 (Liles, 1956, found adults in all 
months; the adult overwinters.). Habitat: in, on, or near bracket fungi, which are the 
sporophores of polyporoid fungi; Ganoderma applanatum (Persoon) is the most 
common host. Life history studies have been published by Liles, 1956, and Pace, 
1967. 
Genus BOLITOPHAGUS Illiger 
corticola (Say). 10 35 53 60 64 103 110 207 210 213 214. April 15 to  September 29. 
Habitat: in woody fungus or under bark of dead trees. 
Genus ELEATES Casey 
depressus (Randall). 207 214. June 28 to July 29. Habitat: in woody fungus or under 
bark of dead trees. 
Tribe RHIPIDANDRINI 
Genus RHIPIDANDRUS LeConte 
paradoxus (Palisot). 25 32. No collecting dates. Habitat: usually in fungi attached to dead 
trees. 
Tribe DIAPERINI 
Genus DIAPERIS Miiller 
maculata Olivier. 10 12 21 24 25 32 33 35 44 45 53 60 64 66 83 84 91 94 100 111 
112 203 207 211 213 214. April 17 to September 26. Habitat: in fungus. 
Genus NEOMIDA Latreille 
bicornis (Fabricius). 10 16 20 21 24 25 33 35 51 64 73 75 80 85 103 110 112 213. 
March 23 to October 23. Habitat: in fungi on old tree stumps. This species and other 
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88 THE GREAT LAKES ENTOMOLOGIST Vol. 6, No. 3 
members of the genus appeared under the generic name Hoplocephala Laporte and 
Brullb until Triplehorn (1965: 374) discovered that Neomida was the older name. 
Genus PLATYDEMA Laporte and Bmllb 
The species of this genus are usually found in fungi, most commonly in fungi on dead 
trees; they are also found under the bark of dead trees. 
americanum Laporte and Brullb. 10 24 25 33 40 47 53 75 110 112 203 207 212 214. 
May 30 to October. 
ellipticum (Fabricius). 24. No collecting date. 
excavatum (Say). 15 21  23 24 25 33 35 43 50 5 1  63 64. April 9 to  October 23. 
picilabrum Melsheimer. 24 25. May 25. 
ruficorne (Sturm). 10 16 2 1  24 25 33 35 36 44 52  65. April 4 to October 23. 
subcostatum Laporte and Brullb. 12 14 21  23 24 25 34 35 36 47 213. May 1 to October 
2. 
teleops Triplehorn. 23 25 33 5 1  64 73. April 4 to September 5. 
Genus SCAPHIDEMA Redtenbacher 
aeneolum (LeConte). 25 207. No collecting dates. Habitat: usually under bark of dead 
trees. 
Genus PENTAPHYLLUS Dejean 
palIidus LeConte. 25 32. July 22 to August. Habitat: usually under bark of dead trees. 
Tribe ULOMINI 
Genus CYNAEUS LeConte 
angustus (LeConte). (Larger black flour beetle.) 21 33 95. May 18 to July 26. This 
species is often a pest of dried stored food products; it is also found under bark of 
dead trees. 
Genus LATHETICUS Waterhouse 
oryzae Waterhouse. (Longheaded flour beetle.) 25. No collecting dates. This species is 
usually a pest of dried stored food products. It is found in many parts of the world. 
Genus TRIBOLIUM MacLeay 
audax Halstead. (American black flour beetle.) 203 207. June 14 to July 7. Habitat: 
often a pest of dried stored food products; also under bark, in leaf mold under logs, 
and in beehives. This species was formerly known as T. madens (Carpentier), but the 
latter does not occur in the United States. All former records of T. madens in North 
America apply to T. audax. 
castaneum (Herbst). (Red flour beetle.) 14 33. February 1 to July 30. Habitat: usually a 
pest of dried stored food products; also under bark of dead trees. It occurs in many 
parts of the world. 
confusum Jacquelin Duval. (Confused flour beetle.) 14 24 25 33 4 1  64. January, April to 
July, October, December. Habitat: usually a pest of dried stored food products; also 
under bark of dead trees. It occurs in many parts of the world. 
Genus DIOEDUS LeConte 
punctatus LeConte. 25. August. Habitat: under bark of dead trees. 
Genus PALORUS Mulsant 
ratzeburgi (Wissmann). (Smalleyed flour beetle.) 70. July. Habitat: usually a pest of dried 
stored food products. It is found in many parts of the world. 
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1973 THE GREAT LAKES ENTOMOLOGIST 
Genus ULOMA Dejean 
The species are usually found under the bark of dead trees. 
imberbis LeConte. 25. April 20 to May 30. 
imprcssa Melsheimer. 21 25 60 66 93 102. April 10 to October. 
mentalis Horn. 24 25 35 60. April 22 to July 23. 
punctulata LeConte. 21 34 35. June 8 to July 22, 
Genus ALPHITOBIUS Dejean 
diaperinus (Panzer). (Lesser mealworm.) 33 73. May 2 to June 27. This species is usually 
a pest of dried stored food products. It is found in many parts of the world. 
Genus CORTICEUS Piller and Mitterpacher 
parallelus (Melsheimer). 47 207 210. June 21. Habitat: under bark of dead trees, usually 
in galleries of Scolytidae. The generic name Hypophloeus Fabricius was used in old 
catalogues. 
Tribe TENEBRIONINI 
Genus CENTRONOPUS Solier 
calcaratus (Fabricius). 10 11 21 23 24 25 35 52 60 64 66 72 102 103 112. June 15 to 
October 11. Habitat: under bark of dead trees. The generic name Scotobates was used 
until recently. 
Genus MERINUS LeConte 
Iaevis (Olivier). 10 20 24 25 33 36 75 207. April 20 to September 21. Habitat: under 
bark of dead trees. 
Genus XYLOPINUS LeConte 
The species are usually found under the bark of dead trees. 
aenescens LeConte. 60. July 7. 
saperdioides (Olivier). 10 21 24 25 32 34 35 50 60 75 83 85 91 103 110 112 207 214. 
April 14 to September 14. 
Genus HAPLANDRUS LeConte 
The species are found under the bark of dead trees. 
concolor LeConte. 75 110 111 112 207 209 211 213 214. May 30 to August 1. 
fulvipes (Herbst). 21 60 207. July 11 to August 11. 
Genus IPHTHINUS Truqui 
opacus LeConte. 66 70 75 93 100 102 111 112 200 205 207 210 211 213 214. April 5 
to September 25. Habitat: under bark of dead trees. 
Genus ALOBATES Motschulsky 
The species are found under bark of dead or dying hardwood trees. 
morio (Fabricius). 24 60 66 90 103 112 207. July 8 to July 26. This species is usually 
known as Alobates barbata (Knoch). Blair (1914:487) showed that its correct name is 
morio. 
pennsylvanicus (DeGeer). 10 12 15 20 21 24 25 31 33 35 41 43 46 51 52 54 61 66 75 
80 81 83 84 90 101 102 103 110 111 112 200 201 207 210 211 213 214. April4 
to November 12. 
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90 THE GREAT LAKES ENTOMOLOGIST Vol. 6, No. 3 
Genus UPIS Fabricius 
ceramboides (Linnaeus). 25 35 53 62 66 75 90 92 93 95 102 103 110 112 200 201 202 
207 209 210 21 1 213 214. May 1 to October 26. Habitat: under bark of dead trees. 
Genus IDIOBATES Casey 
castaneus (Knoch). 10 21 35 52 53. May 23 to September 3. Habitat: under bark of 
dead trees. 
Genus TENEBRIO Linnaeus 
molitor Linnaeus. (Yellow mealworm.) 10 21 24 25 33 34 35 44 45 60 65 66 75 80 83 
85 90 91 102 103 112 207 209 210 211 213. April 15 to August 17. Habitat: 
usually a pest of dried stored food products; also under bark of dead trees. It occurs 
in many parts of the world. 
Genus NEATUS LeConte 
tenebrioides (Palisot). 10 15 16 21 23 24 25 31 32 33 34 35 42 43 45 50 51  52 53 54 
63 64 65 66 74 75 81 84 85 91 93 101 110 112 200 207 210 211 214. April 5 to 
September 1. Habitat: under bark of dead trees. This species has often been included 
in the genus Tenebrio and has often been considered a synonym of N. picipes 
(Herbst), a palearctic species. 
Genus BIUS Mulsant 
estriatus (LeConte). 207. June 29. Habitat: under bark of dead trees. 
Tribe HETEROTARSINI 
Genus PARATENETUS Spinola 
The species are found under bark of dead trees and in plant debris. 
fuscus LeConte. 25 35 64. June to October. 
punctatus Spinola. 25 207. May to July 3. 
Tribe MERACANTHINI 
Genus MERACANTHA Kirby 
contracta (Palisot). 10 16 21 23 24 25 33 34 35 42 44 66 75 91 207 210. May 15 to 
September 21. Habitat: under bark of dead trees. 
Tribe STRONGY LllNI 
Genus STRONGYLIUM Kirby 
The species are found under the bark of dead trees. 
tenuicolle (Say). 10 20 21 23 24 25 33 34 35 44 47 60 64 73 74 85 110 112 207 208 
210. June 9 to August 9. 
terminatum (Say). 21 25 45. July 22. 
APPENDIX 
As stated in the introduction, most of the species in the foregoing list were recorded 
in previous Michigan lists. However, some of those species had been recorded under 
different generic and/or specific names. I have indicated recent synonymies; other 
synonymies will be found in the Leng Catalogue of the Coleoptera of America north of 
Mexico. 
The following species appeared in previous lists but do not appear in my list: 
Blapstinus intewzrptus (Say), of the Pedinini, was recorded by Hubbard and Schwarz 
(1878a and 1878b) and Andrews (1916). I cannot separate this species from B. metallicus 
(F.) in my identifications. A generic revision will undoubtedly resolve this confusion. 
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1973 THE GREAT LAKES ENTOMOLOGIST 91 
Platydema flavipes (Fabricius), of the Diaperini, was recorded by Andrews (1916). 
Platydema laevipes Haldeman, of the Diaperini, was recorded by Andrews (1916). 
Gnatocems maxillosus (Fabricius), of the Ulomini, was recorded by Hubbard and 
Schwarz (1878b). This species is often a pest of dried stored food products; it is found in 
many parts of the world. 
Tenebrio obscurus Fabricius, of the Tenebrionini, was recorded by Hubbard and 
Schwarz (1878b) and Andrews (1916 and 1923). This species is usually a pest of stored 
dried food products; it is found in many parts of the world. 
Paratenetus gibbipennis Motschulsky, of the Heterotarsini, was recorded by Hubbard 
and Schwarz (1878b). 
LITERATURE CITED 
Adams, C. C. 1909. The Coleoptera of Isle Royale, Lake Superior, and their relation to 
the North American centers of dispersal. (p. 157-215, with p. 204-212 by A. B. 
Wolcott) In Adams' An ecological survey of Isle Royale, Lake Superior, 468 p., illus. 
(Rept. Bd. of Geol. Surv. for 1908.) 
Andrews, A. W. 1916. Results of the Mershon Expedition to the Charity Islands, Lake 
Huron: Coleoptera. Mich. Geol. and Biol. Sum., Publ. 20, Biol. Ser. 4:67-108. 
1923. The Coleoptera of the Shiras Expedition to Whitefish Point, 
Chippewa County, Michigan. Pap. Mich. Acad. Sci. Arts and Letters, I, 
1921[1923] :293-309. 
Hatch, M. H. 1924. A list of Coleoptera from Charlevoix County, Michigan. Pap. Mich. 
Acad. Sci. Arts and Letters, 4:543-586. 
Hubbard, H. G., and Schwarz, E. A. 1878a. List of the Coleoptera found in .the Lake 
Superior Region. Proc. Amer. Phil. Soc., 17:627-643. 
1878b. Contribution to a list of the Coleoptera of the Lower Peninsula of 
Michigan. Proc. Amer. Phil. Soc., 17:643-666. 
Liles, M. P. 1956. A study of the life history of the forked fungus beetle, Bolitothems 
cornutus (F'anzer). Ohio J.  Sci., 56(6):329-337, illus. 
Pace, A. E. 1967. Life history and behavior of a fungus beetle, Bolitothems cornutus 
(Tenebrionidae). Univ. Mich. Mus. Zool. Occ. Pap. 653:l-15. 
Townsend, T. 1889. Contribution to a list of the Coleoptera of the Lower Peninsula of 
Michigan. Psyche, 5 :231-325. 
Triplehorn, C. A. 1965. Revision of Diaperini of America north of Mexico with notes on 
extralimital species (Coleoptera: Tenebrionidae). Proc. United States Natl. Mus., 
117:349-458, illus. 
7
Spilman: A List of the Tenebrionidae of Michigan (Coleoptera)
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A List of the Tenebrionidae of Michigan (Coleoptera)

Valparaiso University

The Great Lakes Entomologist Volume 6 Number 3 -- Fall 1973 Number 3 -- Fall 1973 Article 2 August 2017 A List of the Tenebrionidae of Michigan (Coleoptera) T.J. Spilman US Department of Agriculture Follow this and additional works at: https://scholar.valpo.edu/tgle  Part of the Entomology Commons Recommended Citation Spilman, T.J. 2017. "A List of the Tenebrionidae of Michigan (Coleoptera)," The Great Lakes Entomologist, vol 6 (3) Available at: https://scholar.valpo.edu/tgle/vol6/iss3/2 This Peer-Review Article is brought to you for free and open access by the Department of Biology at ValpoScholar. It has been accepted for inclusion in The Great Lakes Entomologist by an authorized administrator of ValpoScholar. For more information, please contact a ValpoScholar staff member at scholar@valpo.edu. 1973 THE GREAT LAKES ENTOMOLOGIST A LIST OF THE TENEBRIONIDAE OF MICHIGAN (COLEOPTERA) T. J. Spilman Entomology Research Division, Agricultural Research Service, United States Departmeni of Agriculture, Washington, D . c . ~  The family Tenebrionidae, the darkling beetles, containing more than 14,000 species, is found primarily in the tropics and arid regions. As would therefore be expected, Michigan's cool, moist environment is not attractive to very many tenebrionids. This list contains 50 identified species and six previously recorded species that were not confirmed by me. Previously 45 species of Tenebrionidae had been recorded in Michigan in various lists. I have not attempted to find all literature records, but I have seen the principal faunal lists of Coleoptera. The three areas of the state, Lower Peninsula, Upper Peninsula, and Isle Royale, are used here to summarize previous lists. In the Lower Peninsula, Hubbard and Schwarz (1878b) recorded 32 species, Townsend (1889) recorded three species, Andrews (1916) recorded 21 species from the Charity Islands (Arenac County) in the mouth of Saginaw Bay, and Hatch (1924) recorded 11 species from Charlevoix County. In the Upper Peninsula, Hubbard and Schwarz (1878a) recorded 16 species from the Lake Superior area, and Andrews (1923) recorded 13 species from Whitefish Point. On Isle Royale, Adams (1909) recorded one species. After duplications in these lists are accounted for, the Lower Peninsula had 34 species, the Upper Peninsula had 25, Isle Royale, of course, had only one, and the whole state had 45. Of the 45 species, six are not recorded in my list; those six are discussed in the Appendix. In my list of 50 identified species, 46 occur in the Lower Peninsula, 26 occur in the Upper Peninsula, none is recorded from Isle Royale, and one species had only a state label on the specimen. This list is based on specimens I have examined from the following institutions: University of Michigan Museum of Zoology, Ann Arbor, lent by Richard D. Alexander; Michigan State University, East Lansing, lent by Roland L. Fischer and the late R. R. Dreisbach; U.S. National Museum of Natural History, Washington, D.C. In addition to !locality records from my identifications, I have included localities published by C. A. Triplehorn in his 1965 revision of the Diaperini. DISTRIBUTIONAL DATA The basic unit of distributional information is the county. Counties are numbered by tiers, beginning in the southwestern corner of the state. The counties in the Lower Peninsula are numbered from 10 to 113, those in the Upper Peninsula from 200 to  214, and Isle Royale 300. The system permits condensation of data and, after a little familiarity, easy visualization of county position in the state by its number alone (see Fig. 1). The numbering of the counties is as follows: 10-Berrien 25-Wayne 44-Shiawassee 11-Cass 30-Allegan 45-Genesee 12-St. Joseph 31-Barry 46-Lapeer 13-Branch 3 2-Eaton 47-St. Clair 14-Hillsdale 33-Ingham 50-Muskegon 15 -Lenawee 34-Livingston 5 1-Montcalm 16-Monroe 35 -Oakland 52-Gratiot 20-Van Buren 36-Macomb 53-Saginaw 21-Kalamazoo 40-Ottawa 54-Tuscola 22-Calhoun 41-Kent 55 -Sanilac 23-Jackson 42-Ionia 60-Oceana 24-Washtenaw 43-Clinton 6 1 -Newaygo l ~ a i l i n ~  address: Department of Entomology, Smithsonian Institution, Washington, D.C. 20560. 1Spilman: A List of the Tenebrionidae of Michigan (Coleoptera)Publ shed by ValpoScholar, 2017THE GREAT LAKES ENTOMOLOGIST Vol. 6, No. 3 Fig. 1. The counties of the State of Michigan indicated by numbers. For explanation, see text page 85. 75 -Arenac 80-Manistee 8 1 -Wexford 82-Missaukee 83-Roscommon 84-Ogemaw 85-IOSCO 90-Benzie 91 -Grand Traverse 92-Kalkaska 2The Great Lakes Entomologist, Vol. 6, No. 3 [2017], Art. 2https://scholar.valpo.edu/tgle/vol6/iss3/2THE GREAT LAKES ENTOMOLOGIST 112-Cheboygan 113 -Presque Isle 200-Gogebic 201 -0ntonagon 202-Houghton 203-Keweenaw Subfamily ASIDINAE Tribe ZOPHERINI Genus PHELLOPSIS LeConte obcordata (Kirby). 112 207 214. July 5 to September 16. Habitat: under bark of dead trees. Subfamily TENEBRIONINAE Tribe PEDININI Genus BLAPSTINUS Dejean The species are often found in debris on ground, also in grasses. Some species in other parts of the United States occasionally build up in great numbers and damage truck crops. metallicus (Fabricius). 10  25 31 35 47 66 83 85 93 95 100 110 213. April 24 to September 3. moestus Melsheimer. 10 25. July 24. Tribe BOLITOPHAGINI Genus BOLITOTHERUS Candkze cornutus (Panzer). 10 20 21 24 25 33 34 35 36 40 44 45 66 75 91 100 110 200 201 204 207 210 21 1 213 214. April 5 to September 26 (Liles, 1956, found adults in all months; the adult overwinters.). Habitat: in, on, or near bracket fungi, which are the sporophores of polyporoid fungi; Ganoderma applanatum (Persoon) is the most common host. Life history studies have been published by Liles, 1956, and Pace, 1967. Genus BOLITOPHAGUS Illiger corticola (Say). 10 35 53 60 64 103 110 207 210 213 214. April 15 to  September 29. Habitat: in woody fungus or under bark of dead trees. Genus ELEATES Casey depressus (Randall). 207 214. June 28 to July 29. Habitat: in woody fungus or under bark of dead trees. Tribe RHIPIDANDRINI Genus RHIPIDANDRUS LeConte paradoxus (Palisot). 25 32. No collecting dates. Habitat: usually in fungi attached to dead trees. Tribe DIAPERINI Genus DIAPERIS Miiller maculata Olivier. 10 12 21 24 25 32 33 35 44 45 53 60 64 66 83 84 91 94 100 111 112 203 207 211 213 214. April 17 to September 26. Habitat: in fungus. Genus NEOMIDA Latreille bicornis (Fabricius). 10 16 20 21 24 25 33 35 51 64 73 75 80 85 103 110 112 213. March 23 to October 23. Habitat: in fungi on old tree stumps. This species and other 3Spilman: A List of the Tenebrionidae of Michigan (Coleoptera)Published by ValpoScholar, 201788 THE GREAT LAKES ENTOMOLOGIST Vol. 6, No. 3 members of the genus appeared under the generic name Hoplocephala Laporte and Brullb until Triplehorn (1965: 374) discovered that Neomida was the older name. Genus PLATYDEMA Laporte and Bmllb The species of this genus are usually found in fungi, most commonly in fungi on dead trees; they are also found under the bark of dead trees. americanum Laporte and Brullb. 10 24 25 33 40 47 53 75 110 112 203 207 212 214. May 30 to October. ellipticum (Fabricius). 24. No collecting date. excavatum (Say). 15 21  23 24 25 33 35 43 50 5 1  63 64. April 9 to  October 23. picilabrum Melsheimer. 24 25. May 25. ruficorne (Sturm). 10 16 2 1  24 25 33 35 36 44 52  65. April 4 to October 23. subcostatum Laporte and Brullb. 12 14 21  23 24 25 34 35 36 47 213. May 1 to October 2. teleops Triplehorn. 23 25 33 5 1  64 73. April 4 to September 5. Genus SCAPHIDEMA Redtenbacher aeneolum (LeConte). 25 207. No collecting dates. Habitat: usually under bark of dead trees. Genus PENTAPHYLLUS Dejean palIidus LeConte. 25 32. July 22 to August. Habitat: usually under bark of dead trees. Tribe ULOMINI Genus CYNAEUS LeConte angustus (LeConte). (Larger black flour beetle.) 21 33 95. May 18 to July 26. This species is often a pest of dried stored food products; it is also found under bark of dead trees. Genus LATHETICUS Waterhouse oryzae Waterhouse. (Longheaded flour beetle.) 25. No collecting dates. This species is usually a pest of dried stored food products. It is found in many parts of the world. Genus TRIBOLIUM MacLeay audax Halstead. (American black flour beetle.) 203 207. June 14 to July 7. Habitat: often a pest of dried stored food products; also under bark, in leaf mold under logs, and in beehives. This species was formerly known as T. madens (Carpentier), but the latter does not occur in the United States. All former records of T. madens in North America apply to T. audax. castaneum (Herbst). (Red flour beetle.) 14 33. February 1 to July 30. Habitat: usually a pest of dried stored food products; also under bark of dead trees. It occurs in many parts of the world. confusum Jacquelin Duval. (Confused flour beetle.) 14 24 25 33 4 1  64. January, April to July, October, December. Habitat: usually a pest of dried stored food products; also under bark of dead trees. It occurs in many parts of the world. Genus DIOEDUS LeConte punctatus LeConte. 25. August. Habitat: under bark of dead trees. Genus PALORUS Mulsant ratzeburgi (Wissmann). (Smalleyed flour beetle.) 70. July. Habitat: usually a pest of dried stored food products. It is found in many parts of the world. 4The Great Lakes Entomologist, Vol. 6, No. 3 [2017], Art. 2https://scholar.valpo.edu/tgle/vol6/iss3/21973 THE GREAT LAKES ENTOMOLOGIST Genus ULOMA Dejean The species are usually found under the bark of dead trees. imberbis LeConte. 25. April 20 to May 30. imprcssa Melsheimer. 21 25 60 66 93 102. April 10 to October. mentalis Horn. 24 25 35 60. April 22 to July 23. punctulata LeConte. 21 34 35. June 8 to July 22, Genus ALPHITOBIUS Dejean diaperinus (Panzer). (Lesser mealworm.) 33 73. May 2 to June 27. This species is usually a pest of dried stored food products. It is found in many parts of the world. Genus CORTICEUS Piller and Mitterpacher parallelus (Melsheimer). 47 207 210. June 21. Habitat: under bark of dead trees, usually in galleries of Scolytidae. The generic name Hypophloeus Fabricius was used in old catalogues. Tribe TENEBRIONINI Genus CENTRONOPUS Solier calcaratus (Fabricius). 10 11 21 23 24 25 35 52 60 64 66 72 102 103 112. June 15 to October 11. Habitat: under bark of dead trees. The generic name Scotobates was used until recently. Genus MERINUS LeConte Iaevis (Olivier). 10 20 24 25 33 36 75 207. April 20 to September 21. Habitat: under bark of dead trees. Genus XYLOPINUS LeConte The species are usually found under the bark of dead trees. aenescens LeConte. 60. July 7. saperdioides (Olivier). 10 21 24 25 32 34 35 50 60 75 83 85 91 103 110 112 207 214. April 14 to September 14. Genus HAPLANDRUS LeConte The species are found under the bark of dead trees. concolor LeConte. 75 110 111 112 207 209 211 213 214. May 30 to August 1. fulvipes (Herbst). 21 60 207. July 11 to August 11. Genus IPHTHINUS Truqui opacus LeConte. 66 70 75 93 100 102 111 112 200 205 207 210 211 213 214. April 5 to September 25. Habitat: under bark of dead trees. Genus ALOBATES Motschulsky The species are found under bark of dead or dying hardwood trees. morio (Fabricius). 24 60 66 90 103 112 207. July 8 to July 26. This species is usually known as Alobates barbata (Knoch). Blair (1914:487) showed that its correct name is morio. pennsylvanicus (DeGeer). 10 12 15 20 21 24 25 31 33 35 41 43 46 51 52 54 61 66 75 80 81 83 84 90 101 102 103 110 111 112 200 201 207 210 211 213 214. April4 to November 12. 5Spilman: A List of the Tenebrionidae of Michigan (Coleoptera)Published by ValpoScholar, 201790 THE GREAT LAKES ENTOMOLOGIST Vol. 6, No. 3 Genus UPIS Fabricius ceramboides (Linnaeus). 25 35 53 62 66 75 90 92 93 95 102 103 110 112 200 201 202 207 209 210 21 1 213 214. May 1 to October 26. Habitat: under bark of dead trees. Genus IDIOBATES Casey castaneus (Knoch). 10 21 35 52 53. May 23 to September 3. Habitat: under bark of dead trees. Genus TENEBRIO Linnaeus molitor Linnaeus. (Yellow mealworm.) 10 21 24 25 33 34 35 44 45 60 65 66 75 80 83 85 90 91 102 103 112 207 209 210 211 213. April 15 to August 17. Habitat: usually a pest of dried stored food products; also under bark of dead trees. It occurs in many parts of the world. Genus NEATUS LeConte tenebrioides (Palisot). 10 15 16 21 23 24 25 31 32 33 34 35 42 43 45 50 51  52 53 54 63 64 65 66 74 75 81 84 85 91 93 101 110 112 200 207 210 211 214. April 5 to September 1. Habitat: under bark of dead trees. This species has often been included in the genus Tenebrio and has often been considered a synonym of N. picipes (Herbst), a palearctic species. Genus BIUS Mulsant estriatus (LeConte). 207. June 29. Habitat: under bark of dead trees. Tribe HETEROTARSINI Genus PARATENETUS Spinola The species are found under bark of dead trees and in plant debris. fuscus LeConte. 25 35 64. June to October. punctatus Spinola. 25 207. May to July 3. Tribe MERACANTHINI Genus MERACANTHA Kirby contracta (Palisot). 10 16 21 23 24 25 33 34 35 42 44 66 75 91 207 210. May 15 to September 21. Habitat: under bark of dead trees. Tribe STRONGY LllNI Genus STRONGYLIUM Kirby The species are found under the bark of dead trees. tenuicolle (Say). 10 20 21 23 24 25 33 34 35 44 47 60 64 73 74 85 110 112 207 208 210. June 9 to August 9. terminatum (Say). 21 25 45. July 22. APPENDIX As stated in the introduction, most of the species in the foregoing list were recorded in previous Michigan lists. However, some of those species had been recorded under different generic and/or specific names. I have indicated recent synonymies; other synonymies will be found in the Leng Catalogue of the Coleoptera of America north of Mexico. The following species appeared in previous lists but do not appear in my list: Blapstinus intewzrptus (Say), of the Pedinini, was recorded by Hubbard and Schwarz (1878a and 1878b) and Andrews (1916). I cannot separate this species from B. metallicus (F.) in my identifications. A generic revision will undoubtedly resolve this confusion. 6The Great Lakes Entomologist, Vol. 6, No. 3 [2017], Art. 2https://scholar.valpo.edu/tgle/vol6/iss3/21973 THE GREAT LAKES ENTOMOLOGIST 91 Platydema flavipes (Fabricius), of the Diaperini, was recorded by Andrews (1916). Platydema laevipes Haldeman, of the Diaperini, was recorded by Andrews (1916). Gnatocems maxillosus (Fabricius), of the Ulomini, was recorded by Hubbard and Schwarz (1878b). This species is often a pest of dried stored food products; it is found in many parts of the world. Tenebrio obscurus Fabricius, of the Tenebrionini, was recorded by Hubbard and Schwarz (1878b) and Andrews (1916 and 1923). This species is usually a pest of stored dried food products; it is found in many parts of the world. Paratenetus gibbipennis Motschulsky, of the Heterotarsini, was recorded by Hubbard and Schwarz (1878b). LITERATURE CITED Adams, C. C. 1909. The Coleoptera of Isle Royale, Lake Superior, and their relation to the North American centers of dispersal. (p. 157-215, with p. 204-212 by A. B. Wolcott) In Adams' An ecological survey of Isle Royale, Lake Superior, 468 p., illus. (Rept. Bd. of Geol. Surv. for 1908.) Andrews, A. W. 1916. Results of the Mershon Expedition to the Charity Islands, Lake Huron: Coleoptera. Mich. Geol. and Biol. Sum., Publ. 20, Biol. Ser. 4:67-108. 1923. The Coleoptera of the Shiras Expedition to Whitefish Point, Chippewa County, Michigan. Pap. Mich. Acad. Sci. Arts and Letters, I, 1921[1923] :293-309. Hatch, M. H. 1924. A list of Coleoptera from Charlevoix County, Michigan. Pap. Mich. Acad. Sci. Arts and Letters, 4:543-586. Hubbard, H. G., and Schwarz, E. A. 1878a. List of the Coleoptera found in .the Lake Superior Region. Proc. Amer. Phil. Soc., 17:627-643. 1878b. Contribution to a list of the Coleoptera of the Lower Peninsula of Michigan. Proc. Amer. Phil. Soc., 17:643-666. Liles, M. P. 1956. A study of the life history of the forked fungus beetle, Bolitothems cornutus (F'anzer). Ohio J.  Sci., 56(6):329-337, illus. Pace, A. E. 1967. Life history and behavior of a fungus beetle, Bolitothems cornutus (Tenebrionidae). Univ. Mich. Mus. Zool. Occ. Pap. 653:l-15. Townsend, T. 1889. Contribution to a list of the Coleoptera of the Lower Peninsula of Michigan. Psyche, 5 :231-325. Triplehorn, C. A. 1965. Revision of Diaperini of America north of Mexico with notes on extralimital species (Coleoptera: Tenebrionidae). Proc. United States Natl. Mus., 117:349-458, illus. 7Spilman: A List of the Tenebrionidae of Michigan (Coleoptera)Published by ValpoScholar, 2017

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A List of the Tenebrionidae of Michigan (Coleoptera)

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