Experiments were conducted in three field environments with the following objectives: (1) to evaluate the progress from seven cycles of reciprocal recurrent selection in Iowa Stiff Stalk Synthetic (BSSS(R)] and Iowa Corn Borer Synthetic #1 (BSCBl(R)] and six cycles of half-sib family selection in \u27Alph\u27 (i.e., BS12) maize (Zea mays L.) populations, (2) to compare the response of unimproved and improved maize variety hybrids to different levels of nitrogen fertilizer and plant density, and (3) to evaluate the influence of nitrogen and plant density on the morphological and physiological traits associated with recurrent selection for grain yield in maize. Each experiment was grown in randomized complete blocks with a split-split-plot arrangement and two replications. Nitrogen fertilizer levels (0, 90, 180, and 270 kg N/ha) were main plots, plant densities (39,500; 59,300; 79,000; and 98,800 plants/ha) were subplots, and five variety hybrids, BSSS(R)CO x BSCBl(R)CO, BSSS(R)C5 x BSCB1(R)C5, BSSS(R)C7 x BSCB1(R)C7, BS12CO x B14A, and BS12C6 x B14A, were randomized as sub-subplots. We obtained data on grain yield and grain-yield components, flowering traits, plant traits, leaf area, leaf orientation, lodging, dry-matter productivity, and harvest index