Se propone una nueva clasificación para Gymnocarpio dryopteridis-Abietion lasiocarpa

Costa, Manuel

Rivas-Martínez, Salvador

Sánchez Mata, Daniel

English

Gestion del Repositorio Documental de la Universidad de Salamanca

GYMNOCARPIO DRYOPTERIDIS-ABIETION LASIOCARPAENOMEN DUBIUM ET NOMEN AMBIGUUMGymnocarpio dryopteridis-Abietion lasiocarpae nomendubium et ambiguumSalvador RIVAS-MARTíNEZ*, Daniel SÁNCHEZ-MATA** & Manuel COSTA**** Phytosociological Research Center, J.M. Usandizaga, 46. E-28409 Los Negrales, Madrid,España. rivas-martinez.cif@tsai.es.** Departamento de Biología Vegetal II (Botánica). Facultad de Farmacia. UniversidadComplutense. E-28040 Madrid, España. dsmata@eucmax.sim.ucm.es.*** Jardín Botánico, Beato Gaspar de Bono, 6. Universidad de Valencia. E-46008 Valencia,España. manuel.costa@uv.esBIBLID [0211 - 9714 (1999) 18, 161- ]Fecha de aceptación de la nota: 30-12-99The alliance Gymnocarpio dryopteridis-Abietion lasiocarpae Wali & Krajina exPeinado, Aguirre & Cruz 1998 (Plant Ecology 137: 192), is supported by the asso-ciation Gymnocarpio dryopteridis-Abietetum lasiocarpae Wali & Krajina ex Pei-nado, Aguirre & Cruz 1998 (l.c.), which has as its nomenclatural type (lectotypus inPEINADO & al., op. cit.: 193) the relevé number 1 included in table 6 of the invalidlypublished name association –(art. 10 of the current issue of the Code of the Phy-tosociological Nomenclature, BARKMAN & al., Vegetatio 67: 145-195. 1986; CPN)–Gymnocarpio (dryopteridis)-Oplopanaco (horridi)-Abieto (lasiocarpae)-Piceetumglaucae Wali & Krajina 1973 (Vegetatio 26(4-6): 258). PEINADO & al. (l.c.) publishno relevés belonging to the new association they validate, nor any relevés of theirsynonymized association, Ptilio (cristae-castrensis)-Gymnocarpio (dryopteridis)-Abieto (lasiocarpae)-Piceetum glaucae Wali & Krajina (l.c.). This situation makes itnecessary to consider the selected relevé (number 1) as the obliged nomenclaturaltype and the protologue of the Wali & Krajina invalid association as the only diag-nostic reference.© Ediciones Universidad de Salamanca Stud. bot. 18, 1999, pp. 161-22ISSN: 0214-3402RIVAS-MARTÍNEZ, SÁNCHEZ-MATA & COSTA (Itinera Geobotanica 12: 56. 1999;30.06.99) propose as nomen dubium (art. 37, 38), the name of the alliance Gym-nocarpio dryopteridis-Abietion lasiocarpae Wali & Krajina ex Peinado & al. 1998(art. 37 CPN) as the supported type-relevé, selected by PEINADO & al. (l.c.), isincompatible with the Wali & Krajina protologue (l.c.). The reason is the abundanceof the characteristic species of the secondary deciduous forests of the order Betulopapyriferae-Populetalia tremuloidis Rivas-Martínez, Sánchez-Mata & Costa 1999such as Alnus viridis (Vill.) Lam. & DC. subsp. sinuata (Regel) A. & D. Löve yBetula papyrifera Marsh., and also the absence of Oplopanax horridus Miq., whichthe Canadian authors (op. cit.: 259) point out in the protologue as the dominantcharacteristic species.We have now also added the article 36 (CPN) in order to reject the PEINADO &al. alliance name due to taxonomic corrections in the characteristic and dominanttrees of this association. On the one hand, PEINADO & al. (l.c.) assure that Wali &Krajina’s Picea glauca really belongs to the hybrid Picea glauca x engelmannii(Picea glauca var. albertiana sensu nobis); on the other hand, SPRIBILLE (Phytocoe-nologia 29(4): 569. 1999) when he refers to the genus Abies species (teste HUNT,Flora of North America 2: 359. 1993) assures that Wali & Krajina’s Abies lasiocarpa(op. cit.: table 6) really belongs to Abies bifolia, and in consequence (art. 43 CPN),he corrects the names of the association and alliance (Gymnocarpio-Abietetum bifo-liae, Gymnocarpio-Abietion bifoliae). No decisions can be made based on the pre-cise geographical location of the places where Wali & Krajina made their relevés, asthe mapped and described territory is so vast (British Columbia: 54º-56ºN/122º-125ºW). It is therefore impossible to decide with certainty as to the taxonomiccorrections on trees belonging to the genus Picea and Abies. We reported in theabove-mentioned vast territory (approx. 42,000 square Km) the presence of Abieslasiocarpa (Hook.) Nutt., Abies bifolia A. Murray bis, Abies lasiocarpa x bifolia,Picea engelmannii Parry ex Engelm., Picea glauca (Moench) Voss and Picea engel-mannii x glauca (Picea glauca var. albertiana (S. Br.) Sarg.). Finally, we would liketo point out that the territory where Wali & Krajina worked belongs to the followingbiogeographical sectors: Canadian Rockies (Canadian Rockies and Omineca Moun-tains subsectors) and Fraser Interior Plateau (Nechako-Fraser Plateau subsector),according to our recent biogeographical proposals for the North American territo-ries (RIVAS-MARTÍNEZ, SÁNCHEZ-MATA & COSTA, op. cit.: 27). As a result, we hereby pro-pose to the Commission of Phytosociological Nomenclature the application of arti-cle 36 (CPN), as the restoration of the original sense of the association will be apermanent source of confusion (nomen ambiguum propositus hoc loco).Nevertheless, it is probable that at least a part of the Wali & Krajina invalidassociation (op. cit.: 258, table 6) can be assigned to the association Piceetumengelmannio-albertianae Rivas-Martínez, Sánchez-Mata & Costa 1999 (Itinera Geo-botanica 12: 152) and thus included in the alliance Piceo engelmannii-Abietionbifoliae Rivas-Martínez, Sánchez-Mata & Costa 1999 (op. cit.: 141), order Pseudot-sugo glaucae-Abietetalia bifoliae Rivas-Martínez, Sánchez-Mata & Costa 1999 (op.cit.: 138), classis Pseudotsugetea glaucae Rivas-Martínez, Sánchez-Mata & Costa1999 (Itinera Geobotanica 13: 350).SALVADOR RIVAS-MARTÍNEZ, DANIEL SÁNCHEZ-MATA & MANUEL COSTAGYMNOCARPIO DRYOPTERIDIS-ABIETION LASIOCARPAE NOMEN DUBIUM ET NOMEN AMBIGUUM162© Ediciones Universidad de Salamanca Stud. bot. 18, 1999, pp. 161-162

Gymnocarpio dryopteridis-Abietion lasiocarpae nomen dubium et nomen ambiguum

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