Peck, Carol J.

Peck, James H.

Taylor, W. Carl

English

UARK (University of Arkansas )

Journal of the Arkansas Academy of Science Volume 41 Article 41 1987 Arkansas Pteridophyte Flora Update: A New Checklist and Additional County-level Occurrence Records James H. Peck University of Arkansas at Little Rock Carol J. Peck University of Arkansas at Pine Bluff W. Carl Taylor Milwaukee Public Museum Follow this and additional works at: https://scholarworks.uark.edu/jaas  Part of the Botany Commons Recommended Citation Peck, James H.; Peck, Carol J.; and Taylor, W. Carl (1987) "Arkansas Pteridophyte Flora Update: A New Checklist and Additional County-level Occurrence Records," Journal of the Arkansas Academy of Science: Vol. 41 , Article 41. Available at: https://scholarworks.uark.edu/jaas/vol41/iss1/41 This article is available for use under the Creative Commons license: Attribution-NoDerivatives 4.0 International (CC BY-ND 4.0). Users are able to read, download, copy, print, distribute, search, link to the full texts of these articles, or use them for any other lawful purpose, without asking prior permission from the publisher or the author. This General Note is brought to you for free and open access by ScholarWorks@UARK. It has been accepted for inclusion in Journal of the Arkansas Academy of Science by an authorized editor of ScholarWorks@UARK. For more information, please contact scholar@uark.edu. Proceedings Arkansas Academy off Science, Vol. 41, 1987 113Arkansas Academy off Science>IThese taxa are part or the Lycopodium inundatum-complex, a group of species plus their hybrids which are cryptic, difficult to tell apartmorphologically (Bruce, 1975; Beitel, 1979; Lellinger, 1985). They maintain their distinctness across their range, in spite of their co-occurrenceas a genus community and their propensity to hybridize. The use of pteridophyte genus communities (communities of many species of one genus)to expand the recognition of taxonomically useful characteristics (such as comparisons ofvegetative and reproductive phenology) forcryptic speciesofpteridophytes was summarized by Wagner and Wagner (1983), who noted that much comparative data which validates the distinctness of speciescan only be ascertained when the taxa co-occur.I Allof these newly reported Lycopodium taxa were found occurring in genus communities across Calhoun Co., at locations where disturbancewas pronounced, including barrow pits, gravel/sand quarries, roadside ditches, cleared pine plantations, and in the midst of refuse of a landfillIoperation. The permanence of these populations is open to study; Amason showed us one location where a large stand onceflurished, but whereonly a few stems were presently evident, apparently having been invaded and replaced by various grasses and sedges. Although presently knownin Arkansas from one county, we fully expect that additional search will establish that they occur throughout the Gulf Coastal Plain in Arkansas.LITERATURE CITED)BEITEL, J. 1979. Clubmosses (Lycopodium) in North America.Fiddlehead Forum 6(5)1-8.LELLINGER, D. B. 1985. A field manual of the ferns and fernallies of the United States and Canada. Smithsonian InstitutionPress, Washington, DC.BRUCE, J. G. 1975. Systematics and morphology of subgenusLepidotus of the genus Lycopodium (Lycopodiaceae). Ph.D. disser-tation, University of Michigan, Ann Arbor, MI. (Univers.SNYDER, L. H., and J. G. BRUCE. 1986. Field guide to the fernsand other pteridophytes of Georgia. University Georgia Press,Athens, GA. 270 pp.Microfilms Intern. 76-9355.)CRANFILL, R. 1980. Ferns and fern allies of Kentucky. Kentucky TAYLOR, W. C. 1984. Arkansas ferns and fern allies. MilwaukeePublic Museum, Milwaukee, WI. 262 pp.Nature Preserves Commission Scientific and Technical Series, No.1. Frankfurt, KY.284 pp.TAYLOR, W. C, and D. DEMAREE. 1979. Annotated listof the fernsand fern allies of Arkansas. Rhodora 81:503-548.CRANFILL, R. 1981. Bog clubmosses (Lycopodiella) in Kentucky.Amer. Fern J. 71:97-100.WAGNER, W. H., and F. S. WAGNER. 1983. Genus communitiesas a systematic tool in the study of New World Botrychium(Ophioglossaceae). Taxon 32:53-62.JOHNSON, R. G., and T. N. McCOY. 1975. Some lycopodiums newto Western Kentucky. Amer. Fern J. 65:29.PECK, JAMES H., CAROL J. PECK, STEVE L. ORZELL, EDWINBRIDGES, and CARL AMASON, Biology Department, University ofArkansas at LittleRock, LittleRock, AR 72204; Department ofNatural Sciences, University of Arkansas at Pine Bluff, Pine Bluff, AR 71601;Texas Natural Heritage Program, Austin, TX 78701; Institute Botanical Research, Austin, TX 78746; Calion, AR 71724.ARKANSAS PTERIDOPHYTE FLORA UPDATE: A NEW CHECKLISTAND ADDITIONALCOUNTY-LEVEL OCCURRENCE RECORDSThis note announces the publication of a new state checklist of pteridophytes (Peck, Peck, and Taylor, 1987) and of a compilation of countyoccurrence record from specimens deposited at the UALR Herbarium (Peck and Peck, 1986). Over the last 152 years, 14 checklists or floras ofthe Arkansas pteridophytes (ferns plus fern allies) were published (Taylor, 1984). Our new checklist of 85 taxa (species plus hybrids) provides atimely update, summarizing the numerous changes in Arkansas pteridophyte floristic information that have developed through the efforts of severalworkers over the last 10 years. As part of his doctoral studies on the Arkansas pteridophyte flora, Taylor (1976) inspected herbarium materialand conducted numerous field trips withD. Demaree to relocate populations of taxa rare in Arkansas (Taylor and Demaree, 1979). Various staffmembers of the Arkansas Natural Heritage Commission and Arkansas Nature Conservancy have continued to relocate rare pteridophyte popula-tions and have located taxa previously unknown to Arkansas (Peck, Orzell, Sundell, and Peck, 1985; Peck, Peacock, and Shepherd, 1985; Peck,Peck, Orzell, Bridges, and Amason, 1987). Over the last six years, two of us (JHP and CJP) have conducted research on the reproductive biologyof disjunct fern populations (Peck, 1985; Peck, 1986; Peck and Peck, 1987) and have encouraged our students to collect ferns, resulting inadditionsof county records from under-collected areas of the state. The summary (Peck and Peck, 1986) of county-occurrence records was compiled froma 1986 inventory of pteridophyte collections deposited at the UALR Herbarium (LRU). A total of 287 new county records belonging to 48 taxawere added to the distributional data presented in the atlas of the Arkansas flora (Smith, 1978). The authors welcome notification ofother additionsto the pteridophyte flora of Arkansas.LITERATURE CITEDPECK, J. H. 1985. Trichomanes petersii (Hymenophyllaceae) in an PECK, J. H., S. L. ORZELL, E. SUNDELL, and C. J. PECK. 1985.Dryopteris ludoviciana and D. Xaustralis new to Arkansas. Amer.Fern J. 75:71.abondoned mine in the Ouachita Mountains ofArkansas. Southw.Nat. 30:463-464."ECK, J. H. 1986. Second locality for Dryopteris carthusiana inArkansas. Amer. Fern J. 76:28-29.PECK, J. H., L. PEACOCK, and W. SHEPHERD. 1985. Observa-tions and new records of ferns naturalizing in Arkansas. Proc. Ark.Acad. Sci. 39:144-145.113Journal of the Arkansas Academy of Science, Vol. 41 [1987], Art. 41Published by Arkansas Academy of Science, 1987114 Proceedings Arkansas Academy of Science, Vol. 41, 1987General NotesPECK, J. H. and C. J. PECK. 1986. Additions to the Arkansaspteridophyte flora. Ark. Nat. Plant Soc. Occ. Pap. No. 7. 4 pp.PECK, J. H.and C. J. PECK. 1987. Field status of Woodsia scopulinain Arkansas. Proc. Ark. Acad. Sci. 41:112.PECK, J. H., C. J. PECK, S. ORZELL, E. BRIDGES, and C.AMASON. 1987. Discovery of Lyco/?ew//M/w-communities on theCoastal Plain of southern Arkansas. Proc. Ark. Acad. Sci.41:112-113.PECK, J. H., C. J. PECK, and W. C. TAYLOR. 1987. Checklistof ferns and fern allies of Arkansas. Ark. Acad. Sci., Ark. BiotaSurv. Checklist No. 47. 4 pp.SMITH, E. B. 1978. An atlas and annotated list of the vascularplants ofArkansas. Dept. Botany and Bacteriology, University ofArkansas, Fayetteville, AR.TAYLOR, W. C. 1976. Arkansas pteridophyta: description anddistribution. Ph.D. dissertation, Southern Illinois University,Carbondale, IL.TAYLOR, W. C. 1984. Arkansas ferns and fern allies. MilwaukeePublic Museum, Milwaukee, WI. 262 pp.TAYLOR, W. C. and D. DEMAREE. 1979. Annotated list of theferns and fern allies of Arkansas. Rhodora 81:503-548.PECK, JAMES H, CAROL J. PECK, and W. CARL TAYLOR, Department ofBiology, University of Arkansas at LittleRock, LittleRock,AR 72204; Department ofNatural Sciences, University of Arkansas at Pine Bluff, Pine Bluff, AR 61901; and Botany Division, Milwaukee PublicMuseum, Milwaukee, WI53233.NEW LOCATIONS FOR PONDBERRY (LINDERA MELISSIFOLIA) INARKANSASPondberry (Lindera melissifolia) belongs to the Laurel Family (Lauraceae) and is closely related to Spicebush (Lindera benzoin). These twoshrubs occupy low-wooded moist habitats withPondberry occurring in water and forming dense vegetative colonies. Pondberry reaches twometersin heighth with little branching whereas Spicebush can attain heights of fivemeters and branches frequently.Before 1985, Pondberry (Lindera melissifolia) was known in Arkansas from a few locations in northern Clay County. One of the originalsites in Arkansas is the Hartwig site in Clay County discovered by Tucker in 1972 (Tucker, 1974a). Pondberry is known from adjacent RipleyCounty in Missouri (Steyermark, 1949).Pondberry is cited by Steyermark (1963) as one ofthe rare shrubs in the United States. This plant is verified as occurring in four to six statesoutside ofArkansas. The U.S. Fish and Wildlife Service classifies Pondberry as a Category 2 species, classified as endangered or threatened, pend-ing further information. Ayensu and DeFilipps (1978) recommended Pondberry for national endangered status, Tucker (1974b) classified Pond-berry as endangered in Arkansas, Berger and Neuner (1979) listed Pondberry as threatened in Louisiana.Pondberry occurs in Arkansas in the Mississippi Embayment Region which has a varied topography ofdepressions and sandy knolls. Saucier(1978) discusses sand dunes and related eolian features of the lower Mississippi River Valley, including such areas in the St. Francis River BasinofArkansas and Missouri. The Arkansas Natural Heritage Commission contracted an aerial survey over this area in March, 1984 to look for poten-tialPondberry sites. The Commission analyzed the results of this survey and determined the depressions with standing water and forest cover asthe best areas to conduct a ground survey.Seventeen such sites were selected as potential Pondberry locations withan additional 12 subdivisions from the original areas in a nine countyarea of Arkansas. The counties include: Clay, Greene, Jackson, Lawrence, Lee, Monroe, Phillips, Randolph and Woodruff. A ground surveywas initiated in the spring of 1985 to search for this plant. Allof the above areas were checked by walking through the sites to determine the presenceor absence of Pondberry. This small shrub was found in seven of the 17 main areas and four of the 12 subdivisions in the above counties. Theresults were positive forone area each for Lawrence and Woodruff Counties and four areas were found with Pondberry in Jackson County. Anadditional site was also found in Clay County. This increases the county distribution from one to four for Lindera melissifolia in Arkansas.Pondberry habitats in Arkansas are threatened by land clearing and many small populations have been damaged or destroyed since 1970(personal communication from the Arkansas Natural Heritage Commission). One site ofhistorical occurrence inClay County for this shrub couldnot be verified in this study. Klomps (1980) mentions the destruction ofan additional Arkansas site where land was converted to row crops. Clear-ing, levelling and destruction of mounds and depressions is currently taking place. Itis hoped with the finding of additional sites during this studythat some areas can be permanently preserved. Thanks are due the Arkansas Natural Heritage Commission for financial support for this research.LITERATURE CITEDAYENSU, E. S. and R. A. DEFILIPPS. 1978. Endangered andThreatened Plants of the United States. Smithsonian Institutionand World Wildlife Fund, Inc., Washington, DC. 301 pp.BERGER, T. J. and M. A. NEUNER. 1979. Directory of StateProtected Species. Association of Systematics Collections.Lawrence, Kansas, n.p.KLOMPS, V. L. 1980. The Status of Lindera melissifolium (Walt.)Blume, Pondberry, in Missouri. Trans. Missouri Acad. Sci. Vol.14:61-66.SAUCIER, R. T. 1978. Sand Dunes and Related Eolian Features ofthe Lower Mississippi River AlluvialValley. Geoscience and Man19:23-40.STEYERMARK, J. A. 1949. Lindera melisaefolia. Rhodora 51:153-162.STEYERMARK, J. A. 1963. Flora of Missouri. Iowa State Univ.Press. 1728 pp.TUCKER, G. E. 1974a. Lindera melissaefolium in Arkansas. Rhodora76:525.TUCKER, G. E. 1974b. Threatened native plants of Arkansas. Pp39-65, in C. T. Crow. 1974. Arkansas natural area plan. State ofArkansas, LittleRock. 248 pp.EDWARD L.RICHARDS, Dept. ofBiological Sciences, Arkansas State University, State University, Arkansas 72467, and STEVE L. ORZELL,Texas Natural Heritage Program, 1700 N. Congress Ave., Austin, Texas 78701.114Journal of the Arkansas Academy of Science, Vol. 41 [1987], Art. 41https://scholarworks.uark.edu/jaas/vol41/iss1/41

Arkansas Pteridophyte Flora Update: A New Checklist and Additional County-level Occurrence Records

ScholarWorks@UARK

Journal of the Arkansas Academy of ScienceVolume 41 Article 411987Arkansas Pteridophyte Flora Update: A NewChecklist and Additional County-level OccurrenceRecordsJames H. PeckUniversity of Arkansas at Little RockCarol J. PeckUniversity of Arkansas at Pine BluffW. Carl TaylorMilwaukee Public MuseumFollow this and additional works at: http://scholarworks.uark.edu/jaasPart of the Botany CommonsThis article is available for use under the Creative Commons license: Attribution-NoDerivatives 4.0 International (CC BY-ND 4.0). Users are able toread, download, copy, print, distribute, search, link to the full texts of these articles, or use them for any other lawful purpose, without asking priorpermission from the publisher or the author.This General Note is brought to you for free and open access by ScholarWorks@UARK. It has been accepted for inclusion in Journal of the ArkansasAcademy of Science by an authorized editor of ScholarWorks@UARK. For more information, please contact scholar@uark.edu, ccmiddle@uark.edu.Recommended CitationPeck, James H.; Peck, Carol J.; and Taylor, W. Carl (1987) "Arkansas Pteridophyte Flora Update: A New Checklist and AdditionalCounty-level Occurrence Records," Journal of the Arkansas Academy of Science: Vol. 41 , Article 41.Available at: http://scholarworks.uark.edu/jaas/vol41/iss1/41Proceedings Arkansas Academy off Science, Vol. 41, 1987 113Arkansas Academy off Science>I These taxa are part or the Lycopodium inundatum-complex, a group of species plus their hybrids which are cryptic, difficult to tell apartmorphologically (Bruce, 1975; Beitel, 1979; Lellinger, 1985). They maintain their distinctness across their range, in spite of their co-occurrenceas a genus community and their propensity to hybridize. The use of pteridophyte genus communities (communities of many species of one genus)to expand the recognition of taxonomically useful characteristics (such as comparisons ofvegetative and reproductive phenology) forcryptic speciesofpteridophytes was summarized by Wagner and Wagner (1983), who noted that much comparative data which validates the distinctness of speciescan only be ascertained when the taxa co-occur.I Allof these newly reported Lycopodium taxa were found occurring in genus communities across Calhoun Co., at locations where disturbancewas pronounced, including barrow pits, gravel/sand quarries, roadside ditches, cleared pine plantations, and in the midst of refuse of a landfillIoperation. The permanence of these populations is open to study; Amason showed us one location where a large stand once flurished, but whereonly a few stems were presently evident, apparently having been invaded and replaced by various grasses and sedges. Although presently knownin Arkansas from one county, we fully expect that additional search will establish that they occur throughout the Gulf Coastal Plain in Arkansas.LITERATURE CITED)BEITEL, J. 1979. Clubmosses (Lycopodium) in North America.Fiddlehead Forum 6(5)1-8.LELLINGER, D. B. 1985. A field manual of the ferns and fernallies of the United States and Canada. Smithsonian InstitutionPress, Washington, DC.BRUCE, J. G. 1975. Systematics and morphology of subgenusLepidotus of the genus Lycopodium (Lycopodiaceae). Ph.D. disser-tation, University of Michigan, Ann Arbor, MI. (Univers.SNYDER, L. H., and J. G. BRUCE. 1986. Field guide to the fernsand other pteridophytes of Georgia. University Georgia Press,Athens, GA. 270 pp.Microfilms Intern. 76-9355.)CRANFILL, R. 1980. Ferns and fern allies of Kentucky. Kentucky TAYLOR, W. C. 1984. Arkansas ferns and fern allies. MilwaukeePublic Museum, Milwaukee, WI. 262 pp.Nature Preserves Commission Scientific and Technical Series, No.1. Frankfurt, KY.284 pp.TAYLOR, W. C, and D. DEMAREE. 1979. Annotated listof the fernsand fern allies of Arkansas. Rhodora 81:503-548.CRANFILL, R. 1981. Bog clubmosses (Lycopodiella) in Kentucky.Amer. Fern J. 71:97-100.WAGNER, W. H., and F. S. WAGNER. 1983. Genus communitiesas a systematic tool in the study of New World Botrychium(Ophioglossaceae). Taxon 32:53-62.JOHNSON, R. G., and T. N. McCOY. 1975. Some lycopodiums newto Western Kentucky. Amer. Fern J. 65:29.PECK, JAMES H., CAROL J. PECK, STEVE L. ORZELL, EDWINBRIDGES, and CARL AMASON, Biology Department, University ofArkansas at LittleRock, LittleRock, AR 72204; Department ofNatural Sciences, University of Arkansas at Pine Bluff, Pine Bluff, AR 71601;Texas Natural Heritage Program, Austin, TX 78701; Institute Botanical Research, Austin, TX 78746; Calion, AR 71724.ARKANSAS PTERIDOPHYTE FLORA UPDATE: A NEW CHECKLISTAND ADDITIONALCOUNTY-LEVEL OCCURRENCE RECORDSThis note announces the publication of a new state checklist of pteridophytes (Peck, Peck, and Taylor, 1987) and of a compilation of countyoccurrence record from specimens deposited at the UALR Herbarium (Peck and Peck, 1986). Over the last 152 years, 14 checklists or floras ofthe Arkansas pteridophytes (ferns plus fern allies) were published (Taylor, 1984). Our new checklist of 85 taxa (species plus hybrids) provides atimely update, summarizing the numerous changes in Arkansas pteridophyte floristic information that have developed through the efforts of severalworkers over the last 10 years. As part of his doctoral studies on the Arkansas pteridophyte flora, Taylor (1976) inspected herbarium materialand conducted numerous field trips withD. Demaree to relocate populations of taxa rare in Arkansas (Taylor and Demaree, 1979). Various staffmembers of the Arkansas Natural Heritage Commission and Arkansas Nature Conservancy have continued to relocate rare pteridophyte popula-tions and have located taxa previously unknown to Arkansas (Peck, Orzell, Sundell, and Peck, 1985; Peck, Peacock, and Shepherd, 1985; Peck,Peck, Orzell, Bridges, and Amason, 1987). Over the last six years, two of us (JHP and CJP) have conducted research on the reproductive biologyof disjunct fern populations (Peck, 1985; Peck, 1986; Peck and Peck, 1987) and have encouraged our students to collect ferns, resulting inadditionsof county records from under-collected areas of the state. The summary (Peck and Peck, 1986) of county-occurrence records was compiled froma 1986 inventory of pteridophyte collections deposited at the UALR Herbarium (LRU). A total of 287 new county records belonging to 48 taxawere added to the distributional data presented in the atlas of the Arkansas flora (Smith, 1978). The authors welcome notification ofother additionsto the pteridophyte flora of Arkansas.LITERATURE CITEDPECK, J. H. 1985. Trichomanes petersii (Hymenophyllaceae) in an PECK, J. H., S. L. ORZELL, E. SUNDELL, and C. J. PECK. 1985.Dryopteris ludoviciana and D. Xaustralis new to Arkansas. Amer.Fern J. 75:71.abondoned mine in the Ouachita Mountains ofArkansas. Southw.Nat. 30:463-464."ECK, J. H. 1986. Second locality for Dryopteris carthusiana inArkansas. Amer. Fern J. 76:28-29.PECK, J. H., L. PEACOCK, and W. SHEPHERD. 1985. Observa-tions and new records of ferns naturalizing in Arkansas. Proc. Ark.Acad. Sci. 39:144-145.113Journal of the Arkansas Academy of Science, Vol. 41 [1987], Art. 41Published by Arkansas Academy of Science, 1987114 Proceedings Arkansas Academy of Science, Vol. 41, 1987General NotesPECK, J. H. and C. J. PECK. 1986. Additions to the Arkansaspteridophyte flora. Ark. Nat. Plant Soc. Occ. Pap. No. 7. 4 pp.PECK, J. H.and C. J. PECK. 1987. Field status of Woodsia scopulinain Arkansas. Proc. Ark. Acad. Sci. 41:112.PECK, J. H., C. J. PECK, S. ORZELL, E. BRIDGES, and C.AMASON. 1987. Discovery of Lyco/?ew//M/w-communities on theCoastal Plain of southern Arkansas. Proc. Ark. Acad. Sci.41:112-113.PECK, J. H., C. J. PECK, and W. C. TAYLOR. 1987. Checklistof ferns and fern allies of Arkansas. Ark. Acad. Sci., Ark. BiotaSurv. Checklist No. 47. 4 pp.SMITH, E. B. 1978. An atlas and annotated list of the vascularplants ofArkansas. Dept. Botany and Bacteriology, University ofArkansas, Fayetteville, AR.TAYLOR, W. C. 1976. Arkansas pteridophyta: description anddistribution. Ph.D. dissertation, Southern Illinois University,Carbondale, IL.TAYLOR, W. C. 1984. Arkansas ferns and fern allies. MilwaukeePublic Museum, Milwaukee, WI. 262 pp.TAYLOR, W. C. and D. DEMAREE. 1979. Annotated list of theferns and fern allies of Arkansas. Rhodora 81:503-548.PECK, JAMES H, CAROL J. PECK, and W. CARL TAYLOR, Department ofBiology, University of Arkansas at LittleRock, LittleRock,AR 72204; Department ofNatural Sciences, University of Arkansas at Pine Bluff, Pine Bluff, AR 61901; and Botany Division, Milwaukee PublicMuseum, Milwaukee, WI53233.NEW LOCATIONS FOR PONDBERRY (LINDERA MELISSIFOLIA) INARKANSASPondberry (Lindera melissifolia) belongs to the Laurel Family (Lauraceae) and is closely related to Spicebush (Lindera benzoin). These twoshrubs occupy low-wooded moist habitats withPondberry occurring in water and forming dense vegetative colonies. Pondberry reaches twometersin heighth with little branching whereas Spicebush can attain heights of fivemeters and branches frequently.Before 1985, Pondberry (Lindera melissifolia) was known in Arkansas from a few locations in northern Clay County. One of the originalsites in Arkansas is the Hartwig site in Clay County discovered by Tucker in 1972 (Tucker, 1974a). Pondberry is known from adjacent RipleyCounty in Missouri (Steyermark, 1949).Pondberry is cited by Steyermark (1963) as one ofthe rare shrubs in the United States. This plant is verified as occurring in four to six statesoutside ofArkansas. The U.S. Fish and Wildlife Service classifies Pondberry as a Category 2 species, classified as endangered or threatened, pend-ing further information. Ayensu and DeFilipps (1978) recommended Pondberry for national endangered status, Tucker (1974b) classified Pond-berry as endangered in Arkansas, Berger and Neuner (1979) listed Pondberry as threatened in Louisiana.Pondberry occurs in Arkansas in the Mississippi Embayment Region which has a varied topography ofdepressions and sandy knolls. Saucier(1978) discusses sand dunes and related eolian features of the lower Mississippi River Valley, including such areas in the St. Francis River BasinofArkansas and Missouri. The Arkansas Natural Heritage Commission contracted an aerial survey over this area in March, 1984 to look for poten-tialPondberry sites. The Commission analyzed the results of this survey and determined the depressions with standing water and forest cover asthe best areas to conduct a ground survey.Seventeen such sites were selected as potential Pondberry locations withan additional 12 subdivisions from the original areas in a nine countyarea of Arkansas. The counties include: Clay, Greene, Jackson, Lawrence, Lee, Monroe, Phillips, Randolph and Woodruff. A ground surveywas initiated in the spring of 1985 to search for this plant. Allof the above areas were checked by walking through the sites to determine the presenceor absence of Pondberry. This small shrub was found in seven of the 17 main areas and four of the 12 subdivisions in the above counties. Theresults were positive forone area each for Lawrence and Woodruff Counties and four areas were found with Pondberry in Jackson County. Anadditional site was also found in Clay County. This increases the county distribution from one to four for Lindera melissifolia in Arkansas.Pondberry habitats in Arkansas are threatened by land clearing and many small populations have been damaged or destroyed since 1970(personal communication from the Arkansas Natural Heritage Commission). One site ofhistorical occurrence inClay County for this shrub couldnot be verified in this study. Klomps (1980) mentions the destruction ofan additional Arkansas site where land was converted to row crops. Clear-ing, levelling and destruction of mounds and depressions is currently taking place. Itis hoped with the finding of additional sites during this studythat some areas can be permanently preserved. Thanks are due the Arkansas Natural Heritage Commission for financial support for this research.LITERATURE CITEDAYENSU, E. S. and R. A. DEFILIPPS. 1978. Endangered andThreatened Plants of the United States. Smithsonian Institutionand World Wildlife Fund, Inc., Washington, DC. 301 pp.BERGER, T. J. and M. A. NEUNER. 1979. Directory of StateProtected Species. Association of Systematics Collections.Lawrence, Kansas, n.p.KLOMPS, V. L. 1980. The Status of Lindera melissifolium (Walt.)Blume, Pondberry, in Missouri. Trans. Missouri Acad. Sci. Vol.14:61-66.SAUCIER, R. T. 1978. Sand Dunes and Related Eolian Features ofthe Lower Mississippi River AlluvialValley. Geoscience and Man19:23-40.STEYERMARK, J. A. 1949. Lindera melisaefolia. Rhodora 51:153-162.STEYERMARK, J. A. 1963. Flora of Missouri. Iowa State Univ.Press. 1728 pp.TUCKER, G. E. 1974a. Lindera melissaefolium in Arkansas. Rhodora76:525.TUCKER, G. E. 1974b. Threatened native plants of Arkansas. Pp39-65, in C. T. Crow. 1974. Arkansas natural area plan. State ofArkansas, LittleRock. 248 pp.EDWARD L.RICHARDS, Dept. ofBiological Sciences, Arkansas State University, State University, Arkansas 72467, and STEVE L. ORZELL,Texas Natural Heritage Program, 1700 N. Congress Ave., Austin, Texas 78701.114Journal of the Arkansas Academy of Science, Vol. 41 [1987], Art. 41http://scholarworks.uark.edu/jaas/vol41/iss1/41

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Arkansas Pteridophyte Flora Update: A New Checklist and Additional County-level Occurrence Records

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