Amason, Carl

Bridges, Edwin

Orzell, Steve L.

Peck, Carol J.

Peck, James H.

English

ScholarWorks@UARK

Journal of the Arkansas Academy of ScienceVolume 41 Article 401987Discovery of Lycopodium Communities in theGulf Coastal Plain Region of ArkansasJames H. PeckUniversity of Arkansas at Little RockCarol J. PeckUniversity of Arkansas at Pine BluffSteve L. OrzellEdwin BridgesBotanical Research Institute Of TexasCarl AmasonFollow this and additional works at: http://scholarworks.uark.edu/jaasPart of the Botany CommonsThis article is available for use under the Creative Commons license: Attribution-NoDerivatives 4.0 International (CC BY-ND 4.0). Users are able toread, download, copy, print, distribute, search, link to the full texts of these articles, or use them for any other lawful purpose, without asking priorpermission from the publisher or the author.This General Note is brought to you for free and open access by ScholarWorks@UARK. It has been accepted for inclusion in Journal of the ArkansasAcademy of Science by an authorized editor of ScholarWorks@UARK. For more information, please contact scholar@uark.edu, ccmiddle@uark.edu.Recommended CitationPeck, James H.; Peck, Carol J.; Orzell, Steve L.; Bridges, Edwin; and Amason, Carl (1987) "Discovery of Lycopodium Communities inthe Gulf Coastal Plain Region of Arkansas," Journal of the Arkansas Academy of Science: Vol. 41 , Article 40.Available at: http://scholarworks.uark.edu/jaas/vol41/iss1/40Proceedings Arkansas Academy of Science, Vol. 41, 1987112General Notes>STATUS REPORT ON THE FERN WOODSIA SCOPULINA INARKANSASWoodsia scopulina D. C. Eat. var. appalachiana (Taylor) Morton (Polypodiophyta: Woodsiaceae) is one of the rarest ferns in the state's |flora, occurring in Arkansas only on Mt. Magazine, Logan Co. (Taylor, 1984; Taylor and Demaree, 1979). The Arkansas population is disjunctover 740 km from the nearest populations to the east in eastern Kentucky and Tennessee (Cranfill, 1980) and to the west in the Rocky Mountains |ofColorado (Lellinger, 198S). The Arkansas plants have a pronounced morphological affinitywithplants in the Southern Appalachian Mountains, \which Taylor (1948) recognized at the varietal rank. In1985, a field study, supported by the Arkansas Nongame Preservation Committee, wasconducted at Mt.Magazine to determine the field status of this fern in Arkansas.The occurrence of W. scopulina was highly restricted, being present ina single location near the summit. The Arkansas population consists'of approximately 700 sporophytes, only 2% of which were juvenile plants; no gametophyte plants were observed. This contrasts markedly with jthe situation for the species in Montana, wherein recruitment was quite conspicuous (Watson and Vazquez, 1981). Although little evidence of suc-cessful sporophyte recruitment from sexual reproduction was observed, the sporophyte plants demonstrated strong vegetative vigor (# fronds perplant) and strong reproductive vitality (# spores per plant).Approximately 80% of the population occurs on the cliff-face, with the remainder within 10 m on the talus below. Inboth habitats, over I90% of the sporophytes were reproductively fertile (producing spores). Sporophytes on the cliff-face averaged 18 fronds per plant, while plants fon the talus averaged 1 1 fronds per plant. Sporophytes on the cliff-face averaged 11 fertile fronds, while plants ofthe talus averaged 7 fertile frondsper plant. Cliff-face plants averaged 674 sori per frond, while talus plants averaged 439 sori per frond.These and other reproductive and ecologic differences between plants on the cliff-face and those on the talus result in a disproportionatecontribution by cliff-face plants to the total annual spore production of this population. The entire population produces an estimated 22 billionspores each year, of which 20 billionare produced bycliff-face plants. Their 10:1 advantage inspore production is greater than their 4: 1 advantagein number ofplants. For this reason, maintenance ofplants on the cliff-face may be an extremely important factor to consider inthe developmentof any management plan or recovery plan that is dedicated to the protection of this population.In that the present population seems to have minimal recruitment from sexual reproduction, even though itis most certainly saturating thelocal environment withspores, the population should be considered extremely fragile. Based on the line ofinquiry presented here, we suggest thatany future forest management or recreational development plans of the summit of Mt. Magazine should preclude alterations in the vicinity ofthis population.LITERATURE CITEDBROWN, D. F. M. 1964. A monographic study of the fern genusWoodsia. Nova. Hedw. Beih. 16:1-154.CRANFILL, R. 1980. Ferns and fern allies of Kentucky. KentuckyNature Preserves Commission Scientific and Technical Series No.1. Frankfurt, KY.LELLINGER,D. B. 1985. A field manual of the ferns and fern alliesof the United States and Canada. Smithsonian Institution Press,Washington, DC.MORTON, C. V. 1950. Notes on the ferns of the eastern UnitedStates. Amer. Fern J. 57:104-106.TAYLOR, T. M.C. 1947. New species and combinations in Woodsia,section Perrinia. Amer. Fern J. 37:84-88.TAYLOR, W. C. 1984. Arkansas ferns and fern allies. MilwaukeePublic Museum, Milwaukee, WI. 262 pp.TAYLOR, W. C. and D. DEMAREE. 1979. Annotated list of theferns and fern allies of Arkansas. Rhodora 81:503-548.WATSON, P. J., and M.VAZQUEZ. 1981. Comparative ecology ofWoodsia scopulina sporophytes and gametophytes. Amer. FernJ. 71:3-9.PECK, JAMESH. and CAROLJ. PECK, Department ofBiology, University ofArkansas at LittleRock, LittleRock, AR 72204, and DepartmentofNatural Sciences, University ofArkansas at Pine Bluff,Pine Bluff, AR 71601.DISCOVERY OF LYCOPODIUM COMMUNITIES INTHE GULF COASTAL PLAINREGION OF ARKANSASUntilrecently, the pteridophytes in the genus Lycopodium were represented in Arkansas by two species (Taylor, 1984; Taylor and Demaree,1979). A few, small populations of the Shinin Clubmoss (L. lucidulum Michx.) occur in the Ozark Region, representing extreme southwesterndisjuncts of this species. Widely dispersed populations of the Appressed Clubmoss (L.appressum [Chapm.] Lloyd&Underw.) occur in the GulfCoastal Plain Region, representing extreme northwestern populations ofthis species. Over the last three collecting seasons (1985-1987), the authorshave added appreciably to the floristic knowledge about Lycopodium in Arkansas. Amason noted the widespread occurrence and large localabundance of Lycopodium in Calhoun Co., AR. Orzell and Bridges detected that the populations were mixed, and included taxa not previouslyreported from Arkansas. Peck and Peck expanded the field survey, added additional populations with unreported taxa, and compared Arkansasmaterial with that of plants from across Southeastern United States held at Missouri Botanical Garden (MO).These efforts resulted in the discovery of additional populations of L. appressum in Calhoun and Garland counties, previously thought tobe quite restricted in its Arkansas occurrence. We report as new to Arkansas three clubmoss species: L. carolinianum L., L. alopecuroides L.,and L.prostratum Harper. We also report as new to Arkansas three clubmoss hybrids: L. X bruceii Lellinger (L. appressum XL. prostratum),L. X copelandii (L. alopecuroides X L. appresum), and L. alopecuroides X L.prostratum.These Lycopodium taxa range along the Atlantic Coastal Plain from Florida to Massachusetts and along the GulfCoastal Plain from Floridawestward to Texas (Bruce, 1965; Synder and Bruce, 1986). They have the ability to survive, at least for a while, in interior continental areas farnorthward from the Gulf of Mexico, in that they were located in southern Kentucky (Johnson and McCoy, 1975; Cranfill, 1980; 1981).112Journal of the Arkansas Academy of Science, Vol. 41 [1987], Art. 40Published by Arkansas Academy of Science, 1987Proceedings Arkansas Academy off Science, Vol. 41, 1987 113Arkansas Academy off Science>IThese taxa are part or the Lycopodium inundatum-complex, a group of species plus their hybrids which are cryptic, difficult to tell apartmorphologically (Bruce, 1975; Beitel, 1979; Lellinger, 1985). They maintain their distinctness across their range, in spite of their co-occurrenceas a genus community and their propensity to hybridize. The use of pteridophyte genus communities (communities of many species of one genus)to expand the recognition of taxonomically useful characteristics (such as comparisons ofvegetative and reproductive phenology) forcryptic speciesofpteridophytes was summarized by Wagner and Wagner (1983), who noted that much comparative data which validates the distinctness of speciescan only be ascertained when the taxa co-occur.I Allof these newly reported Lycopodium taxa were found occurring in genus communities across Calhoun Co., at locations where disturbancewas pronounced, including barrow pits, gravel/sand quarries, roadside ditches, cleared pine plantations, and in the midst of refuse of a landfillIoperation. The permanence of these populations is open to study; Amason showed us one location where a large stand onceflurished, but whereonly a few stems were presently evident, apparently having been invaded and replaced by various grasses and sedges. Although presently knownin Arkansas from one county, we fully expect that additional search will establish that they occur throughout the Gulf Coastal Plain in Arkansas.LITERATURE CITED)BEITEL, J. 1979. Clubmosses (Lycopodium) in North America.Fiddlehead Forum 6(5)1-8.LELLINGER, D. B. 1985. A field manual of the ferns and fernallies of the United States and Canada. Smithsonian InstitutionPress, Washington, DC.BRUCE, J. G. 1975. Systematics and morphology of subgenusLepidotus of the genus Lycopodium (Lycopodiaceae). Ph.D. disser-tation, University of Michigan, Ann Arbor, MI. (Univers.SNYDER, L. H., and J. G. BRUCE. 1986. Field guide to the fernsand other pteridophytes of Georgia. University Georgia Press,Athens, GA. 270 pp.Microfilms Intern. 76-9355.)CRANFILL, R. 1980. Ferns and fern allies of Kentucky. Kentucky TAYLOR, W. C. 1984. Arkansas ferns and fern allies. MilwaukeePublic Museum, Milwaukee, WI. 262 pp.Nature Preserves Commission Scientific and Technical Series, No.1. Frankfurt, KY.284 pp.TAYLOR, W. C, and D. DEMAREE. 1979. Annotated listof the fernsand fern allies of Arkansas. Rhodora 81:503-548.CRANFILL, R. 1981. Bog clubmosses (Lycopodiella) in Kentucky.Amer. Fern J. 71:97-100.WAGNER, W. H., and F. S. WAGNER. 1983. Genus communitiesas a systematic tool in the study of New World Botrychium(Ophioglossaceae). Taxon 32:53-62.JOHNSON, R. G., and T. N. McCOY. 1975. Some lycopodiums newto Western Kentucky. Amer. Fern J. 65:29.PECK, JAMES H., CAROL J. PECK, STEVE L. ORZELL, EDWINBRIDGES, and CARL AMASON, Biology Department, University ofArkansas at LittleRock, LittleRock, AR 72204; Department ofNatural Sciences, University of Arkansas at Pine Bluff, Pine Bluff, AR 71601;Texas Natural Heritage Program, Austin, TX 78701; Institute Botanical Research, Austin, TX 78746; Calion, AR 71724.ARKANSAS PTERIDOPHYTE FLORA UPDATE: A NEW CHECKLISTAND ADDITIONALCOUNTY-LEVEL OCCURRENCE RECORDSThis note announces the publication of a new state checklist of pteridophytes (Peck, Peck, and Taylor, 1987) and of a compilation of countyoccurrence record from specimens deposited at the UALR Herbarium (Peck and Peck, 1986). Over the last 152 years, 14 checklists or floras ofthe Arkansas pteridophytes (ferns plus fern allies) were published (Taylor, 1984). Our new checklist of 85 taxa (species plus hybrids) provides atimely update, summarizing the numerous changes in Arkansas pteridophyte floristic information that have developed through the efforts of severalworkers over the last 10 years. As part of his doctoral studies on the Arkansas pteridophyte flora, Taylor (1976) inspected herbarium materialand conducted numerous field trips withD. Demaree to relocate populations of taxa rare in Arkansas (Taylor and Demaree, 1979). Various staffmembers of the Arkansas Natural Heritage Commission and Arkansas Nature Conservancy have continued to relocate rare pteridophyte popula-tions and have located taxa previously unknown to Arkansas (Peck, Orzell, Sundell, and Peck, 1985; Peck, Peacock, and Shepherd, 1985; Peck,Peck, Orzell, Bridges, and Amason, 1987). Over the last six years, two of us (JHP and CJP) have conducted research on the reproductive biologyof disjunct fern populations (Peck, 1985; Peck, 1986; Peck and Peck, 1987) and have encouraged our students to collect ferns, resulting inadditionsof county records from under-collected areas of the state. The summary (Peck and Peck, 1986) of county-occurrence records was compiled froma 1986 inventory of pteridophyte collections deposited at the UALR Herbarium (LRU). A total of 287 new county records belonging to 48 taxawere added to the distributional data presented in the atlas of the Arkansas flora (Smith, 1978). The authors welcome notification ofother additionsto the pteridophyte flora of Arkansas.LITERATURE CITEDPECK, J. H. 1985. Trichomanes petersii (Hymenophyllaceae) in an PECK, J. H., S. L. ORZELL, E. SUNDELL, and C. J. PECK. 1985.Dryopteris ludoviciana and D. Xaustralis new to Arkansas. Amer.Fern J. 75:71.abondoned mine in the Ouachita Mountains ofArkansas. Southw.Nat. 30:463-464."ECK, J. H. 1986. Second locality for Dryopteris carthusiana inArkansas. Amer. Fern J. 76:28-29.PECK, J. H., L. PEACOCK, and W. SHEPHERD. 1985. Observa-tions and new records of ferns naturalizing in Arkansas. Proc. Ark.Acad. Sci. 39:144-145.113Journal of the Arkansas Academy of Science, Vol. 41 [1987], Art. 40http://scholarworks.uark.edu/jaas/vol41/iss1/40

Discovery of Lycopodium Communities in the Gulf Coastal Plain Region of Arkansas

UARK (University of Arkansas )

Journal of the Arkansas Academy of Science Volume 41 Article 40 1987 Discovery of Lycopodium Communities in the Gulf Coastal Plain Region of Arkansas James H. Peck University of Arkansas at Little Rock Carol J. Peck University of Arkansas at Pine Bluff Steve L. Orzell Edwin Bridges Botanical Research Institute Of Texas Carl Amason Follow this and additional works at: https://scholarworks.uark.edu/jaas  Part of the Botany Commons Recommended Citation Peck, James H.; Peck, Carol J.; Orzell, Steve L.; Bridges, Edwin; and Amason, Carl (1987) "Discovery of Lycopodium Communities in the Gulf Coastal Plain Region of Arkansas," Journal of the Arkansas Academy of Science: Vol. 41 , Article 40. Available at: https://scholarworks.uark.edu/jaas/vol41/iss1/40 This article is available for use under the Creative Commons license: Attribution-NoDerivatives 4.0 International (CC BY-ND 4.0). Users are able to read, download, copy, print, distribute, search, link to the full texts of these articles, or use them for any other lawful purpose, without asking prior permission from the publisher or the author. This General Note is brought to you for free and open access by ScholarWorks@UARK. It has been accepted for inclusion in Journal of the Arkansas Academy of Science by an authorized editor of ScholarWorks@UARK. For more information, please contact scholar@uark.edu. Proceedings Arkansas Academy of Science, Vol. 41, 1987112General Notes>STATUS REPORT ON THE FERN WOODSIA SCOPULINA INARKANSASWoodsia scopulina D. C. Eat. var. appalachiana (Taylor) Morton (Polypodiophyta: Woodsiaceae) is one of the rarest ferns in the state's |flora, occurring in Arkansas only on Mt. Magazine, Logan Co. (Taylor, 1984; Taylor and Demaree, 1979). The Arkansas population is disjunctover 740 km from the nearest populations to the east in eastern Kentucky and Tennessee (Cranfill, 1980) and to the west in the Rocky Mountains |ofColorado (Lellinger, 198S). The Arkansas plants have a pronounced morphological affinitywithplants in the Southern Appalachian Mountains, \which Taylor (1948) recognized at the varietal rank. In1985, a field study, supported by the Arkansas Nongame Preservation Committee, wasconducted at Mt.Magazine to determine the field status of this fern in Arkansas.The occurrence of W. scopulina was highly restricted, being present ina single location near the summit. The Arkansas population consists'of approximately 700 sporophytes, only 2% of which were juvenile plants; no gametophyte plants were observed. This contrasts markedly with jthe situation for the species in Montana, wherein recruitment was quite conspicuous (Watson and Vazquez, 1981). Although little evidence of suc-cessful sporophyte recruitment from sexual reproduction was observed, the sporophyte plants demonstrated strong vegetative vigor (# fronds perplant) and strong reproductive vitality (# spores per plant).Approximately 80% of the population occurs on the cliff-face, with the remainder within 10 m on the talus below. Inboth habitats, over I90% of the sporophytes were reproductively fertile (producing spores). Sporophytes on the cliff-face averaged 18 fronds per plant, while plants fon the talus averaged 1 1 fronds per plant. Sporophytes on the cliff-face averaged 11 fertile fronds, while plants ofthe talus averaged 7 fertile frondsper plant. Cliff-face plants averaged 674 sori per frond, while talus plants averaged 439 sori per frond.These and other reproductive and ecologic differences between plants on the cliff-face and those on the talus result in a disproportionatecontribution by cliff-face plants to the total annual spore production of this population. The entire population produces an estimated 22 billionspores each year, of which 20 billionare produced bycliff-face plants. Their 10:1 advantage inspore production is greater than their 4: 1 advantagein number ofplants. For this reason, maintenance ofplants on the cliff-face may be an extremely important factor to consider inthe developmentof any management plan or recovery plan that is dedicated to the protection of this population.In that the present population seems to have minimal recruitment from sexual reproduction, even though itis most certainly saturating thelocal environment withspores, the population should be considered extremely fragile. Based on the line ofinquiry presented here, we suggest thatany future forest management or recreational development plans of the summit of Mt. Magazine should preclude alterations in the vicinity ofthis population.LITERATURE CITEDBROWN, D. F. M. 1964. A monographic study of the fern genusWoodsia. Nova. Hedw. Beih. 16:1-154.CRANFILL, R. 1980. Ferns and fern allies of Kentucky. KentuckyNature Preserves Commission Scientific and Technical Series No.1. Frankfurt, KY.LELLINGER,D. B. 1985. A field manual of the ferns and fern alliesof the United States and Canada. Smithsonian Institution Press,Washington, DC.MORTON, C. V. 1950. Notes on the ferns of the eastern UnitedStates. Amer. Fern J. 57:104-106.TAYLOR, T. M.C. 1947. New species and combinations in Woodsia,section Perrinia. Amer. Fern J. 37:84-88.TAYLOR, W. C. 1984. Arkansas ferns and fern allies. MilwaukeePublic Museum, Milwaukee, WI. 262 pp.TAYLOR, W. C. and D. DEMAREE. 1979. Annotated list of theferns and fern allies of Arkansas. Rhodora 81:503-548.WATSON, P. J., and M.VAZQUEZ. 1981. Comparative ecology ofWoodsia scopulina sporophytes and gametophytes. Amer. FernJ. 71:3-9.PECK, JAMESH. and CAROLJ. PECK, Department ofBiology, University ofArkansas at LittleRock, LittleRock, AR 72204, and DepartmentofNatural Sciences, University ofArkansas at Pine Bluff,Pine Bluff, AR 71601.DISCOVERY OF LYCOPODIUM COMMUNITIES INTHE GULF COASTAL PLAINREGION OF ARKANSASUntilrecently, the pteridophytes in the genus Lycopodium were represented in Arkansas by two species (Taylor, 1984; Taylor and Demaree,1979). A few, small populations of the Shinin Clubmoss (L. lucidulum Michx.) occur in the Ozark Region, representing extreme southwesterndisjuncts of this species. Widely dispersed populations of the Appressed Clubmoss (L.appressum [Chapm.] Lloyd&Underw.) occur in the GulfCoastal Plain Region, representing extreme northwestern populations ofthis species. Over the last three collecting seasons (1985-1987), the authorshave added appreciably to the floristic knowledge about Lycopodium in Arkansas. Amason noted the widespread occurrence and large localabundance of Lycopodium in Calhoun Co., AR. Orzell and Bridges detected that the populations were mixed, and included taxa not previouslyreported from Arkansas. Peck and Peck expanded the field survey, added additional populations with unreported taxa, and compared Arkansasmaterial with that of plants from across Southeastern United States held at Missouri Botanical Garden (MO).These efforts resulted in the discovery of additional populations of L. appressum in Calhoun and Garland counties, previously thought tobe quite restricted in its Arkansas occurrence. We report as new to Arkansas three clubmoss species: L. carolinianum L., L. alopecuroides L.,and L.prostratum Harper. We also report as new to Arkansas three clubmoss hybrids: L. X bruceii Lellinger (L. appressum XL. prostratum),L. X copelandii (L. alopecuroides X L. appresum), and L. alopecuroides X L.prostratum.These Lycopodium taxa range along the Atlantic Coastal Plain from Florida to Massachusetts and along the GulfCoastal Plain from Floridawestward to Texas (Bruce, 1965; Synder and Bruce, 1986). They have the ability to survive, at least for a while, in interior continental areas farnorthward from the Gulf of Mexico, in that they were located in southern Kentucky (Johnson and McCoy, 1975; Cranfill, 1980; 1981).112Journal of the Arkansas Academy of Science, Vol. 41 [1987], Art. 40Published by Arkansas Academy of Science, 1987Proceedings Arkansas Academy off Science, Vol. 41, 1987 113Arkansas Academy off Science>IThese taxa are part or the Lycopodium inundatum-complex, a group of species plus their hybrids which are cryptic, difficult to tell apartmorphologically (Bruce, 1975; Beitel, 1979; Lellinger, 1985). They maintain their distinctness across their range, in spite of their co-occurrenceas a genus community and their propensity to hybridize. The use of pteridophyte genus communities (communities of many species of one genus)to expand the recognition of taxonomically useful characteristics (such as comparisons ofvegetative and reproductive phenology) forcryptic speciesofpteridophytes was summarized by Wagner and Wagner (1983), who noted that much comparative data which validates the distinctness of speciescan only be ascertained when the taxa co-occur.I Allof these newly reported Lycopodium taxa were found occurring in genus communities across Calhoun Co., at locations where disturbancewas pronounced, including barrow pits, gravel/sand quarries, roadside ditches, cleared pine plantations, and in the midst of refuse of a landfillIoperation. The permanence of these populations is open to study; Amason showed us one location where a large stand onceflurished, but whereonly a few stems were presently evident, apparently having been invaded and replaced by various grasses and sedges. Although presently knownin Arkansas from one county, we fully expect that additional search will establish that they occur throughout the Gulf Coastal Plain in Arkansas.LITERATURE CITED)BEITEL, J. 1979. Clubmosses (Lycopodium) in North America.Fiddlehead Forum 6(5)1-8.LELLINGER, D. B. 1985. A field manual of the ferns and fernallies of the United States and Canada. Smithsonian InstitutionPress, Washington, DC.BRUCE, J. G. 1975. Systematics and morphology of subgenusLepidotus of the genus Lycopodium (Lycopodiaceae). Ph.D. disser-tation, University of Michigan, Ann Arbor, MI. (Univers.SNYDER, L. H., and J. G. BRUCE. 1986. Field guide to the fernsand other pteridophytes of Georgia. University Georgia Press,Athens, GA. 270 pp.Microfilms Intern. 76-9355.)CRANFILL, R. 1980. Ferns and fern allies of Kentucky. Kentucky TAYLOR, W. C. 1984. Arkansas ferns and fern allies. MilwaukeePublic Museum, Milwaukee, WI. 262 pp.Nature Preserves Commission Scientific and Technical Series, No.1. Frankfurt, KY.284 pp.TAYLOR, W. C, and D. DEMAREE. 1979. Annotated listof the fernsand fern allies of Arkansas. Rhodora 81:503-548.CRANFILL, R. 1981. Bog clubmosses (Lycopodiella) in Kentucky.Amer. Fern J. 71:97-100.WAGNER, W. H., and F. S. WAGNER. 1983. Genus communitiesas a systematic tool in the study of New World Botrychium(Ophioglossaceae). Taxon 32:53-62.JOHNSON, R. G., and T. N. McCOY. 1975. Some lycopodiums newto Western Kentucky. Amer. Fern J. 65:29.PECK, JAMES H., CAROL J. PECK, STEVE L. ORZELL, EDWINBRIDGES, and CARL AMASON, Biology Department, University ofArkansas at LittleRock, LittleRock, AR 72204; Department ofNatural Sciences, University of Arkansas at Pine Bluff, Pine Bluff, AR 71601;Texas Natural Heritage Program, Austin, TX 78701; Institute Botanical Research, Austin, TX 78746; Calion, AR 71724.ARKANSAS PTERIDOPHYTE FLORA UPDATE: A NEW CHECKLISTAND ADDITIONALCOUNTY-LEVEL OCCURRENCE RECORDSThis note announces the publication of a new state checklist of pteridophytes (Peck, Peck, and Taylor, 1987) and of a compilation of countyoccurrence record from specimens deposited at the UALR Herbarium (Peck and Peck, 1986). Over the last 152 years, 14 checklists or floras ofthe Arkansas pteridophytes (ferns plus fern allies) were published (Taylor, 1984). Our new checklist of 85 taxa (species plus hybrids) provides atimely update, summarizing the numerous changes in Arkansas pteridophyte floristic information that have developed through the efforts of severalworkers over the last 10 years. As part of his doctoral studies on the Arkansas pteridophyte flora, Taylor (1976) inspected herbarium materialand conducted numerous field trips withD. Demaree to relocate populations of taxa rare in Arkansas (Taylor and Demaree, 1979). Various staffmembers of the Arkansas Natural Heritage Commission and Arkansas Nature Conservancy have continued to relocate rare pteridophyte popula-tions and have located taxa previously unknown to Arkansas (Peck, Orzell, Sundell, and Peck, 1985; Peck, Peacock, and Shepherd, 1985; Peck,Peck, Orzell, Bridges, and Amason, 1987). Over the last six years, two of us (JHP and CJP) have conducted research on the reproductive biologyof disjunct fern populations (Peck, 1985; Peck, 1986; Peck and Peck, 1987) and have encouraged our students to collect ferns, resulting inadditionsof county records from under-collected areas of the state. The summary (Peck and Peck, 1986) of county-occurrence records was compiled froma 1986 inventory of pteridophyte collections deposited at the UALR Herbarium (LRU). A total of 287 new county records belonging to 48 taxawere added to the distributional data presented in the atlas of the Arkansas flora (Smith, 1978). The authors welcome notification ofother additionsto the pteridophyte flora of Arkansas.LITERATURE CITEDPECK, J. H. 1985. Trichomanes petersii (Hymenophyllaceae) in an PECK, J. H., S. L. ORZELL, E. SUNDELL, and C. J. PECK. 1985.Dryopteris ludoviciana and D. Xaustralis new to Arkansas. Amer.Fern J. 75:71.abondoned mine in the Ouachita Mountains ofArkansas. Southw.Nat. 30:463-464."ECK, J. H. 1986. Second locality for Dryopteris carthusiana inArkansas. Amer. Fern J. 76:28-29.PECK, J. H., L. PEACOCK, and W. SHEPHERD. 1985. Observa-tions and new records of ferns naturalizing in Arkansas. Proc. Ark.Acad. Sci. 39:144-145.113Journal of the Arkansas Academy of Science, Vol. 41 [1987], Art. 40https://scholarworks.uark.edu/jaas/vol41/iss1/40

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Discovery of Lycopodium Communities in the Gulf Coastal Plain Region of Arkansas

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