Male Uca tangeri place mudballs in the area surrounding their burrows each
low tide to delineate territory boundaries. We investigated the sequence of mudball
placement by noting the distance at which each mudball was placed and its location in
one of eight sectors around the burrow. Ten of the 13 focal males placed their mudballs
in a pattern that was significantly different from random. No relationship was found
between mudball distance from burrow and order of placement in the mudball sequence.
However, most males placed each mudball in the same or a nearby sector to that of the
previous mudball