Nowak et al.1 wish to explain why the version of kin selection theory that is summarised by the formula R>c/b (c=cost of performing 'altruistic' act, b=benefit derived by recipient of act, R=relatedness between the two) is of little utility for understanding the evolution of eusociality. But in trying to do so they omit much that is relevant and risk misrepresenting the issue to anyone who is not familiar with the literature. A fairer account would include the following facts
