The homolid spermatozoon, as exemplified by #Homola sp., #Paromola sp. and #Paromola petterdi, differs markedly from spermatozoa of crabs of the #Heterotremata - #Thoracotremata assemblage but agrees with the sperm of dromiids, in the strongly anteroposteriorly depressed acrosome (apomorphy?) and the capitate form of the perforatorium (a major synapomorphy seen nowhere else in the #Crustacea). These similarities support inclusion of the #Dromiidae and #Homolidae in a single grouping, the #Podotremata$. The homolid perforatorium differs from that of dromiids in the autapomorphic spiked-wheel form the anterior expansion. (D'après résumé d'auteur
