Razowski, J.

Revistes Catalanes amb Accés Obert

Miscel.lania Zoologica 20.1 (1997) 125 
Generic composition of the 
New World Archipini (Lepidoptera, 
Tortricidae) with description of 
two new genera and two new 
species 
J. Razowski 
Razowski, J , 1997 Generic composition of the New World Archipini (Lepidoptera, Tortricidae) with 
description of two new genera and two new species MISC Zool, 20 1 125-130 
Gener~c ornpos~t~on f the New World Archlplni (Lepldoptera, Tortrlcldae) wlth descrlptlon o f  two 
new genera and two new spec~es - The geographic distribution of the genera of Archipini from the 
New World is discussed and compared with that of other regions The 11 genera found in the 
Nearctic subregion are al1 known frorn the Palaearctic subregion In the Neotropical region there 
are six genera only, three being endernic Two genera, viz, Tacertaenia with T polonorurn and 
Sychnovalva with S syrrhapta are described as new 
Key words Lepidoptera, Tortricidae, Archipini, New World 
(Rebut: 24 V 96; Acceptació cond~cional: 16 XI 96; Acc. definitiva: 28 1 97) 
J Razowskl, lnst o f  Systernatlcs and Evolut~on o f  An~rnals, Pol~sh Acaderny o f  Sc~ences, 31-016 
Kraków, Slawkowska 17, Poland 
lntroduction 
The generic composition o f  the tortricid 
tribes o f  the New World has not  been dis- 
cussed previously. Moreover, data on the 
Neotropical faunas are s t i l l  very incomplete. 
The tribe Archipini has a world-wide dis- 
tribution and the majority of genera and 
species occur in the Oriental region and the 
Palaearctic subregion. In the Palaearctic 
subregion which is the best studied area, 
25 genera occur of which 12 are shared 
wi th the Oriental region. 
The aim o f  this work is t o  present a 
preliminary summary o f  the generic com- 
position o f  the tribe Archipini in the New 
World. Although a comparison wi th that o f  
other tribes is at present impossible, it i s  
clear that  the Archipini are poorly repre- 
sented in  the Neotropical region. 
ISSN 021 1-6529 O 1997 Museu de Zoologia l 
- -- 
- 
126 Razowski 
Material and methods gion, not  expanding far south (except for 
one widely distributed species recorded in  
The present data are based on the material the south from California and Florida, and 
deposited in the largest entomological in- central Mexico). 
stitutions, which are mainly American, and The genus Pandemis is abundant in spe- 
in some private collections. cies in temperate and tropical Asia as well 
as in the Afrotropical region where it oc- 
curs in Madagascar and South Africa. The 
Distribution of the genera of Archipini Nearctic species reaches as far south as Cali- 
in the New World fornia. 
Two genera, Argyrotaenia, Stepyhens, 
The Archipini are represented by only 15 1852 and Clepsis, occur over a wide area 
genera in the New World. Only nine genera o f  the New World, the former f rom north- 
are recorded from the Nearctic subregion. ern Canada t o  Argentina and the latter t o  
Cudonigera Obraztsov & Powell, 1977 is Peru. Argyrotaenia is represented in  the 
endemic in this area. This monotypical genus Holarctic region by a single very widely 
is very close to  Choristioneura Hübner, 1825 distributed Palaearctic species and by over 
differing only in one character of the fe- 35 Nearctic species. A similar number o f  
male genitalia. species occurs in  the Neotropical region. 
Two genera, viz., Batodes Guenée,1845 There are only slight differences between 
and probably Cacoecimorpha Obraztsov, the species and only a few South Ameri- 
1954 were artificially introduced into North can species are better differentiated, par- 
America. ticularly concerning the  genitalia. The 
Four genera (Archips Hübner, 1825, number o f  species o f  Clepsis increases 
Pandemis Hübner, 1825, Adoxophyes Mey- slightly f rom Canada t o  Mexico, and in  
rick, 1881 and Clepsis Guenée, 1845) are South America the genus is represented 
widely distributed in the Palaearctic sub- by five species in  Peru, Venezuela and 
region and the Oriental region. The distri- Guatemala and five other species in  Co- 
bution o f  Adoxophyes is even wider as there lombia. On basis o f  unexamined material 
are several species in the Australian region the author assumes that  this genus i s  more 
and in North America there are t w o  species abundant in species, specially in the moun- 
distributed in its northern parts. tains o f  Peru and Colombia. Three o f  the 
The genera Syndemis Hübner, 1825 and species closely related w i th  C. smicrotes 
Aphelia Hübner, 1825 are Holarctic in  dis- (Walsingham, 1914) occur in  Canada and 
tribution; the  former i s  represented in  14 in  Mexico. Al l  Central American and 
the Palaearctic subregion by a single spe- South American species belong t o  this 
cies, and the latter by numerous species group. Several species described or placed 
and al1 subgenera. The Nearctic species of in  Smicrotes Clemens, 1860 have been 
Aphelia Hübner belong t o  t w o  subgenera; incorrectly included in  P~choloma Ste- 
one species i s  included i n  Zolotherses phens, 1829 which is exclusively a Palaearctic 
Lederer, 1859 four in Aphelia s. str. They genus. Of the remaining species o f  Clepsis 
are widely distributed throughout North one i s  o f  Holarctic distribution, and six 
America f rom Alaska and Quebec t o  Texas. are Nearctic, al1 occurring in  the northern 
Archips Hübner i s  known from al1 of and central areas of North America and 
Asia and in  North America i s  represented more southwards in the mountains (e.g. 
by 21 species distributed mainly in the north- in  California). 
ern and central parts of the subregion but Another Nearctic genus, Durangarchips 
reaching southwards as far as California, Powell, 1991 is monotypical and known 
Texas and Florida. only from Durango (Mexico). 
Choristoneura has a similar distribution In the Neotropical region, apart from 
but  does not  penetrate into the Oriental the last two  genera, occurs ldolatteria Wal- 
region. Seventeen species occur in North singham, 1913 wi th eight species distrib- 
America and are known mainly from the uted between Guatemala and Bolivia and 
northern and central parts of the subre- two  genera described below, al1 most prob- 
Miscel.lania Zoologica 20.1 (1997) 
ably endemic in this region. They are de- 
scribed from Southern Brazil, but their range 
may be much larger. 
As i t  has not been possible to  examine 
any specimens of Nesochoris Clarke, 1965 
the systematic position of this genus can- 
not yet be discussed. 
The tribe Archipini, seems to  be repre- 
sented in the New World by a rather small 
number of the genera and species. Some 
endemic small genera may yet be discov- 
ered in very little studied territories, spe- 
cially in the mountains of the western part 
of the Neotropical region. Many new spe- 
cies may be added to  Argyrotaenia and 
Clepsis. More comprehensive collecting may 
add many taxa as i t  did, for instance, in 
Mexico. 
Descriptions of new taxa 
Tacertaenia n. gen. 
Type-species: Tacertaenia polonorum n. sp. 
Externally similar t o  the representatives 
of Argyrotaenia Stephens. Venation and 
sexual dimorphism as in the mentioned 
genus. 
Male genitalia (figs 1-3) 
Tegumen very broad, with small, narrow 
ends of pedunculi; uncus very large, bifid 
postmedially, with rounded apical parts; 
socius small, membranous, hairy; gnathos 
arms broad, with terminal parts plate- 
shaped, apically rounded, connected with 
one another by means of broad membrane, 
without medial plate; vinculum small, as in 
Argyrotaenia; valva elongate, with dorso- 
terminal part expanding slightly dorsally, 
more strongly sclerotized than the remain- 
ing costal area, sacculus as long as valva, 
angulate at its end, extending dorsally 
along its caudal edge, provided with minute 
free termination, fold distinct, pulvinus ill- 
defined; transtilla band-shaped, broaden- 
ing slightly basally; juxta ovate; aedeagus 
slender, curved, with short coecum penis, 
cornuti missing. 
Female genitalia (fig. 4) 
Eighth tergite and papilla analis large, slen- 
der, apophyses moderate; sterigma rather 
broad in median part, with slender lateral 
arms well sclerotized along anterior edges, 
ostium bursae very small; colliculum small, 
membranous; ductus bursae rather short 
with ductus seminalis extending post-medi- 
ally; signum absent. 
Remarks 
Supposedly closest to  Argyrotaenia as the 
shape of forewings, coloration and sexual 
dimorphism show. The genitalia are similar 
only in the shape of transtilla, aedeagus 
and bursa copulatrix which, however, is 
variable in Argyrotaenia (the sclerite of the 
basal part of ductus bursae is often miss- 
ing). The supposed autapomorphies are 
the broad, rounded apically dista1 parts of 
the uncus, the arms of gnathos connected 
by means of a membrane, the atrophy of 
median, terminal part of the gnathos, the 
enlarged, somewhat stronger sclerotized 
dorsal termination of the valva. The ab- 
sence of the cornuti and the signum are 
convergent reductions often observed in 
this subfamily. The new genus i s  also close 
t o  the new genus described below although 
i t  differs in the absence of median plate of 
the gnathos and its lateral processes, and 
the absence of the signum. 
Tacertaenia polonorum n. sp. 
Male 
Alar expanse 13-15 mm; labial palpus as 
long as diameter of eye, ferruginous to  rust 
yellowish, head, thorax and forewing pat- 
tern concolorous; the latter consisting of 
dorso-basal suffusion, median fascia inter- 
rupted subcostally and subtriangular, large 
subapical blotch; ground-colour much paler 
and creamer, sprinkled rust; fringes in torna1 
half concolorous with ground-colour, other- 
wise with pattern. Hindwing dirty cream, 
tinged grey in anal area, with yellow in 
remaining periphery; fringes concolourous 
with adjacent parts of wing. Variation: pat- 
tern occasionally brown-grey, more or less 
distinct, ground-colour at dorsum pale. Fe- 
male (17-19 mm) almost unicolorous rust, or 
with weak darker pattern before apex and 
tornus; hindwing orange-ferruginous, paler 
basally, brown-grey in anal area. 
l 
128 Razowski 
Figs. 1-7. Male and female genitalia. 1-4. Tacertaenia polonorum n. sp.: 1-3. cf paratype.; 
4. 9 paratype. 5-7. Sychnovalva syrrhapta n. sp.: 5,6. cf holotype; 7. 9 paratype. 
Genitalia del macho y de la hembra. 1-4. Tacertaenia polonorum sp. n.: 1-3. cf 
paratipo; 4. 9 paratipo. 5-7. Sychnovalva syrrhapta sp. n.: 5,6. 8 holotipo;7. 9 paratipo. 
Miscel.lania Zoologica 20.1 (1997) 129 
Genitalia 
As described for the genus. 
sentatives of Archipini (e.g. lsodemis 
Diakonoff, 1952, Homona Walker,1853) in 
having large, membranousfolds of the valva. 
Material studied However, i t  differs from them in the transtilla 
Holotype, 'Brazylia, Santa Catarina, Rio which suggests its affinity to  the group of 
Vermelho, 968 m, 21.11.1973, A. & J. Ra- genera allied with Clepsis. The supposed 
zowski'; coll. V.O. Becker, Planaltina, Brazil. 
Paratypes: 37 males and 17 females from 
type-locality, dated 19-22 1 1  1973 and 
6 111 1973, Rio Negro, Paraná, 900 m., 9 11 1973 
and 14 km NW Sao Bento do Sul, 850 m, 1 
and 23 11 1973. 
Note 
This species is described in honour of the 
Polish speaking inhabitants of the villages 
autapomorphies of this genus are the curved 
base of the transtilla with its proximal proc- 
ess and the pencil hair of the dorsal end of 
the valva. 
Genitalia 
As described for the genus. 
Distribution 
Brazil: Santa Catarina, Paraná. 
in Paraná and Santa Catarina, descendants 
of the Polish immigrants, who helped me so 
much in collecting the moths. Sychnovalva syrrhapta n. sp. 
Alar expanse 15 mm; labial palpus longer 
Sychnovalva n. gen. than diameter of eye; head and tegula 
greyish white; costal part of termen not 
Type-species Sychnovalva syrrhapta n. sp. oblique. Ground-colour yellowish ochreous, 
strigulated and sprinkled brownish; remain- 
Forewing rather broad; costal fold very slen- ders of median fascia and subapical blotch 
der reaching beyond mid-costa; apex very brownish; terminal area of wing veins suf- 
short; termen straight beyond apex. fused brownish; fringes brownish cream. 
Hindwing cream, suffused brownish yellow 
Male genitalia (figs 5,6) on periphery, fringes similar. 
Tegumen small; uncus long, provided with 
ventral brush; socius very small, membra- Genitalia 
nous; gnathos arms slender, terminal plate As described above. 
small; valva large with membranous disc, 
fold large; a pencil of long hair at dorso- Material studied 
apical part of valva; sacculus without free Holotype, male: 'Brazylia, Santa Catarina, 
termination; base of transtilla with curved, Rio Vermelho, 968 m, 26 11 1973, A. & J. 
dentate apical lobe from middle of which Razowski'; coll. V. O. Becker, Planaltina, 
a submembranous process extends; ae- Brasil. Paratype, female similarly labelled 
deagus slender with coecum penis curved but dated 20 11 1973. 
proximally and caulis long; several cornuti 
in vesica. 
Acknowledgements 
Female genitalia (fig. 7) 
Papilla analis small; apophyses very slender; The author is grateful to two anonymous 
sterigma arms small, broad basally; colli- reviewers mainly for the linguistic corrections. 
culum large, rather membranous wi th  
submedian sclerite and anterior lobe; cestum 
reaching 213 of length of ductus bursae, Resumen 
broadening in Corpus bursae; this latter 
minutely spined; signum absent. Composición genérica de los Archipini del 
Nuevo Mundo (Lepidoptera, Tortricidae) con 
Remarks descripción de cuatro nuevos géneros y dos 
This genus resembles some Oriental repre- nuevas especies 
130 Razowski 
Se discute y compara la distribución subregión Paleártica. En la reg ión 
geográfica de los géneros Archipini del Neotropical sólo hay seis géneros, tres de 
Nuevo Mundo con los de otras regiones. ellos endémicos. Se describen dos nuevos 
Los 11 géneros encontrados en la géneros, viz. Tacertaenia con T: polonorum 
subregión Neártica son conocidos de la y Sychonovalva con S. syrrhapta. 


Generic composition of the New World Archipini (Lepidoptera, Torticidae) with description of two new genera and two new species

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