STÉBING, O., PERIS, J.B. &amp; FIGUEROLA, R. 1993. Sideritis molinae un nuevo endemismo ibérico. Bot. Complutensis IB: 113-121  Se describe una nueva especie del género Sideritis L. para la Flora Ibérica

Figuerola, R.

Peris, Juan Bautista

Stübing, G.

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Sideritis molinae a new Iberian endemie
O. STDBING, LB. PERIS & R. FIGUEROLA
Departamento de Biología Vegetal. U.I. Fitografía, Facultad de Farmacia,
Universidad de Valencia, Spain.
Resumen:
STÉBING, O., PERIS, J.B. & FIGUEROLA, R. 1993. Sideritis molinae un nuevo endemismo
ibérico. Bot. Complutensis IB: 113-121
Se describe una nueva especie del género Sideritis L. para la Flora Ibérica.
Abstract:
STCBING, O., PERIS, J.B. & FICUEROLA, R. 1993. Sideritis molinae a new Iberian endemic.
Bot. Complutensis 18: 113-121
A description is given of a new species of the genus Sideritis L. for Ihe Spanish Flora.
INTRODUCTION
After the revision of the subsection Gymnocarpae Font Quer of the
genus Sideritis L. in the western part of the Mediterranean region (PERIS
& al. 1990), we are now working on the subsection Carpostegiatae Font
Quer (FIGUEROLA & al. 1991). As a consequence of studying abundant
herbarium material we have found a plant that in our opinion must be des-
cribed as a new endemie species for the Spanish Flora. Ihe plant has been
included within Sideritis endressi described from the Pyrennees, despite
the existence of significant morphological and chorological differences
between both taxa.
Betónica Complutensis 18: 113-121. Edil. Universidad Complutense 1993
114 O. StiSbing, J. B. Peris & R. Figuerola
DESCRIPTION AND TYPIFICATION
Sideritis mo/mac Stiibing, Peris & Figuerola sp. nova
Sideritis endressii Willk. fma. laxespicata Degen & Debeaux in 1-ter-
vier, Ruil. Acad. mt. Géogr. Rot. (Le Mons) 17: 196 (1907).
Sideritis endressii Willk. subsp. laxespicata (Degen & Debeaux) Hey-
wood, Rol. Jour. Linn. Soc. (London) 65: 355 (1972).
Descriplion: Planta fruticosa, robusta, 40-60 cm longa, ramosa e sua
base. Síipites juvenes castaneae-viridis coloris, quadranguli, cum fasejis
colenquimae maxime apparentibus, hirsuti in basi, cum pilis articularibus
pluricellularibus (3-5 cellulae) 1 .5-2.5 mm, erecti ve] leviter flexi. Juve-
num stipitum pars superior est sub-hirsuta cum pilis pluricellularibus (3-4
cellulae) flexis ad inferioren partem et longis 0.05-1 mm. Folia basalia ex
parte infima 0.6-0.7 x 0.2 cm longa, oblonga-lanceolata, crenato-ciliata et
subhirsuta. Folia media el superiora 3 x 1-2 cm longa, oblonga-lanceolata,
cum dentibus formata, usque a fol jis subacutis ad rotunda. Inflorescentia
in spicae forma cum 5-9(-10) verticilastris, generatim distantes ac cum 8-
10 floribus quisque. Folia floralia 2-3 x 2-3 cm, cauces leviter aequantia
vel prominentia, cordata-ovata, viridia-pallida, hirsuta cum pilis plurice-
llularibus (1-3 cellulae) cum principalibus nerviis et secundiis visilibus
copiosisque in spicae forma (18-24) subaequalibus et ciliatis. Calix 1.4-
1.6 cm, hirsutus cum pilis pluricellularibus longis atque raris pilis glandu-
laribus brevitus, cum nerviis visibilibus, cum dentibus erectis marginis
ciliati el apice obtuso cum acumine. Corolla lutea pallida, 1 .6-2 cm.
Habitat in sub-baeticis montibus provinciae Jaen (in silivis Cazorlae,
silvis Poti, silvis Sccurae atque finibus Albacete inter 600-1700 m supra
solos calcareos).
Dedicata in memoriam Professoris Andreae Molina, amici et colegae, a
vila nuper discess¡.
Description: Perennial, 40-60 cm, branched from the base, young stems
brown-greenish, 4-angled, with collenchyma ribs, hirsute al the base with
articulate 3- to 5-celled trichomes, erect or slightly deflected and 1.5-2.5
mm long. Upper pan of te yomxg stems subhirsute, with defleaed 3- lo
4-celled hairs 00.5-1 mm long. Basal leaves 0.6-0.7 x 0.2 cm, oblong-lan-
ceolale, crenale-ciliale and subhirsute. Middle and upper leaves 3 x 1-2
cm, oblong-lanceolate, subacute to rounded, mucronate. serrate. hirsute.
wiíh 2- to 3-celled trichomes. Inflorescence spikelike, wilh 5-9(-10) vertí.
cillasíers, generally disíant and with 8-10 flowers. Bracís 2-3 x. 2-3 cm,
equaling or silightly surpassing the calyx, cordate-ovate, green-yellowish,
hirsute with 1- lo 3-celled trichomes, with 8-JO main aud secondary veins
visible and 18-24 subequal ciliate spinescent teeth. Calyx 1.4-1.6 cm, hir-
sute, with long pluricellular hairs and a few short glandular hairs; veins
Sideriris molinae a new Iberian endemic 115
Fg. 1. Localion of Sideritis molinar.
116 0. Stñbing, J. fi. Peris & R. Figuerola
MF
A
Fg. 2. Siderftis molinae.
Sideritis molinae a new Iberian endenjie
lo mm
we
=
Ña
10 mm
10 e
“1
Fig. 3. Basal leal: Kl 5. mal/nec, A-2 5. endressii. Sten, leal: 8< 5. mcI/nec B-2S endresgil. Erare
C-l 5. molinee, C-2 8. endressii. Calyx with corolla: D-I 8. mal/nec, D-2 8. cndrcssii. Stem: E-! 5. mcli-
“ar, E-2 S endresgí. (Material usad br drawíngs 5. molinar: MA 45030!, 8. cndrcssii: BCF 35040).
117
1.0 mm
II
w
e,e,
Ña
10 mm
‘.0 10 mm

Sideritis molinee a new Iberian endemír 119
visible; teelh erect and ciliate and apex blunt buí with a mucro. Corolla
1.6-2 cm long, yelowish.
Typification: We follow the same indications as in PERIs & al. (1990).
Type material (1-folotype). Fig. 4.
A.MA 480301. (Specimen in te middle.)
a U 3622/Herbario C. Soriano. Macizo de Segura-Cazorla/ISideritis
endressii Willk./subsp. laxespicata (Degen & Debeaux)//Jaén, Quesada:
alrededores de Puerto Lorente,/3OSWGOOS9, 1700 m.//5-VII-1975//Leg.:
González Rebollar, Muñoz Garmendia & Soriano/Dei.: C. Soriano
Table 1. Comparison beíween Sideritis molinae and Sideritis endressu.
RELATION TO OTHER TAXA
Sideriris mo/mac has been subordinated to Sideritis endressii Willk.,
first as a form and later a subspecies, on ihe basis of te crenate leaves.
The attached table and figures show clearly the differences between both
taxa, which are reinforced by the different distribution.
MATERIAL EXAMINED
Sideritis mo/mac: Jaén: quesada, alrededores de Puerto Lorente,
308WGOOS9, 1700, 5-7-1975, González Rebollar & al. (MA 480301); La
S. molinae .5. endressii
Plant erect-ascendent cred
Young stems 1-2 mm diameter subhirsute 2-3 mm diameter hirsute
Inflorescence 7-10 verticilíasters distant 5-7 verticillasters crowded
with 4-10 flowers with 10-12 flowers
Bracts 2-3 x 2-3cm 0.8-1 x 1-1.5cm
18-24 teeth 12-14 teeth
Calyx hirsute bit-sote
1.4-1.6cm 0.9-1 cm
teeth erect tceth patent
Corolla yellowish yellowish
1.6-2cm 1.2-1.4cm
Distriburion Subbetic mountamns Pyrenees
O. Stñbing, .1. fi. Peris & R. Figuercia120
8-
1,
e
o
~ II As•~4
844n14., V&nId Wn.
k.~r’,•4. ,,.,.,d,~’k. Sn
Hg.5.SidcritiS cndn’5Si.
Sideriris molinaea new Iberian endemir 121
Iruela, Valle del Guadalquivir junto a la pista del Cortijo del Cantalar,
3OSWHO7OI, 800 m, 19-6-75, González Rebollar & al. (MA 480302); La
Iruela, junto a la pista del Zarzaral, 3OSWHO5OI, 1220 m, 22-6-1975,
González Rebollar & al. (MA 480304); Pontones, Barranco del Arroyo
Montero, 30SWH2121, 650 m, 24-7-75, González Rebollar & al. (MA
480303); Cazorla, Barranco del Arroyo de la Mesa, margen izquierda,
30SWG0595, 1000 m, lapiaz de calizas microcristalinas, 28-5-1976, Gon-
zález Rebollar & al. (MA 480305).
Sideritis endrcssii: Rosellón: Declives pedregosos esquistosos junto a
la ermita de la Consolació, sobre Cotíliure. Lugar clásico, 2-5-36, Font
Quer (BCF 3540).
ACKNOWLEDGMENTS
We should like to thank Amparo Olivares for the drawings
REFERENCES
FIOUERoLA, R., STUBING, O. & PERIs. JE. (1991). Nomenclature and typification of 5. angushi-
folia Lag. and 5. tragcriganum Lag. Taxon 40:123-129.
FONT QIJER, P. (1924). Estudis sobre morfologia ¡ nomenclatura de les Sideritis. Trab«¡os del
Museo de Ciencias Naturales de Barcelona, 5 (4): 3-35.
PERIS, iB., STÚBING, O. & FIGUEROLA, R. (1990). An outline revision of the subsection Cym-
nocerpar Font Quer of the genus Sideritis L. (Lamiaccae) ¡o the western part of the Medí-
terranean region. ficlaniral Joernal cfthcLinnean Sor/ay, 103: 1-37.
Recibido 23 dc septiembre de 1992
Aceptado /2 de febrero de 1992


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