16p. Incluye suplemento que contiene: Appendix 2. Species examined. Taxon, origin and voucher information. [http://www.ingentaconnect.com/content/iapt/tax/2011/00000060/00000002/art00020/supp-data/11677lopezsuppl][EN] We investigated the phylogeny of the Arctioid lineage of the Arctium-Cousinia complex in an attempt to clarify the
conflictive generic boundaries of Arctium and Cousinia. The study was based on analyses of one nuclear (ITS) and two chloroplastic
(trnL-trnT-rps4, rpl32-trnL) DNA regions of 37 species and was complemented with morphological evidence where
possible. Based on the results, a broadly redefined monophyletic genus Arctium is proposed. The subgenera Hypacanthodes and
Cynaroides are not monophyletic and are suppressed. In contrast, the traditional sectional classification of the genus Cousinia
is maintained. The genera Anura, Hypacanthium and Schmalhausenia are reduced to sectional level.Peer reviewe
