Whitehead, Paul F.

Biblioteca Digital de les Illes Balears

Observations on Coleoptera 
of Mallorca, Balearic Islands 
(amendments) 
Paul F. WHITEHEAD 
SHNB Whitehead, P,F. 1995. Observations on COleoptera of Mallorca, Balearic Islands (amendments). Bo". Soco Hist. Nat. Balears, 38:163-165. ISSN 0212-260X. Palma de Mallorca. 
~ Diverse ecological and taxonomical aspects 01 some Coleoptera, which appeared in a previous article, are discussed and commented on. Keywords: Coleoptera, Mallorca, taxomony. 
SOCIET AT D'HISTÓRIA 
NATURAL DE LES BAlEARS 
OBSERVACIONS SOBRE COLEOPTERS DE MALLORCA, ILLES BALEARS 
(PUNTUALlTZACIONS). Es discuteixen i comenten diferents aspectes ecologics 
i taxonomics d'alguns coleopters tractats en un anterior article. 
Paraules e/au: Coleoptera, Mallorca, taxonomia. 
Paul F. WHITEHEAD, Moor Leys, Little Comberton, Pershore, Worcestershire, 
WR10 3EH, England. 
Recepció del manuscrit: 5-set-95; revisió acceptada: 24-oct-95. 
Introduction 
My earlier paper in this Bulletin 
(Whitehead, 1993) discussed a number 
of significant Coleoptera from Mallorca, 
with a strong emphasis on the conser-
vation of species from the crucially 
important north coast wetlands and 
littoral. They include species of Cara-
bidae and Hydrophilidae characteristic of 
productive, regularly inundated fen, an 
extensive prime habitat which contri-
butes to the magnificent varied scenery 
of the area. 
A number of misprints crept into 
my earlier paper through no fault of my 
own, which I am now amending, as well 
as updating knowledge of particular 
species. 
Staphylinidae 
The genus Sepedophilus Gistel. 
Mention was made of two species 
on page 49, both having now been 
studied by Dr. M. Schülke, the leading 
authority of the group. The record of 
Sepedophilus pedicularius (Gravenhorst) 
(Lago Esperanza, Port d'Alcúdia, 
25.10.1990) must be deleted. The spe-
cimen is a female, and according to Dr. 
Schülke cannot therefore be determined, 
but an affinity with Sepedophilus 
164 BolI. Soco Hist. Nat. Ba/ears, 38 (1995) 
FIg. 1. a) Kemerton, Worcestershire, 
England, 7.8.1992. b) Alcanada, Mallorca, 
Balearic Islands, April 1987. The general 
morphological conformation and microsculptu-
re of the Balearic insect agrees with S. 
testaceus, but turther work on the 
morphology of the genital internal armature 
of a larger sample may, in the author's view, 
provide a basis tor taxonomic revision. 
Fig. 1. a) Kemerton, Worcestershire, Angla-
terra, 7.8.1992. b) Alcanada, Mal/orca, /l/es 
Balears, abril 1987. La conformació morfoló-
gica general així com la microescultura de 
/'insecte balear coincideixen amb els de S. 
testaceus, peró estudis més precisos sobre 
la morfologia de /'armadura genital interna a 
un exemplar més gros podria, segons 
/'autor, donar l/oc a una revisió taxonómica. 
obtusus (Luz e) (Central and southern 
Europe, near East, certainly not Iberian) 
is apparent. 
Dr. Schülke has confirmed my 
identification of Sepedophilus testaceus 
(Fabricius) (coast near lila d'Alcanada, 
April 1987) commenting that the 
structural details of the aedeagus are 
somewhat atypical. Fig. 1 a shows the 
internal structure of an aedeagus of a 
male S. testaceus from Kemerton, 
Worcestershire, England (7.8.1992, at 
Perenniporia fraxinea (Fr.) Ryvarden on 
Fraxinus exce/sior Unnaeus), of fairly 
typical appearance (Hammond, 1973). 
The internal structure of the aedeagus 
of the Alcanada specimen is shown in 
Fig. 1 b. In England, this species is 
characteristic of fungoid angiosperms. 
Its occurrence on Mallorca on Pinus 
ha/epensis Miller, is more reminiscent of 
Sepedophilus /usitanicus Hammond. 
Pselaphldae 
The entry on page 50 should have 
read as follows: "Brachyg/uta g/obuli-
collis aubéii. Tournier. 80 in flood debris 
on beach by Torrent de S'Albufereta, 
21-25.10.1990; Brachyg/uta schüppelii 
Aubé, a few, Parc de S'Albufera, 
23.10.1990. Examples of both species 
have been seen by Dr. C. Besuchet, 
Geneva". 
Scarabaeldae 
Martín-Piera & Coca-Abia (1992) 
have shown that Rhizotrogus /epidus 
Schaufuss is an insular form of the 
dominantly Iberian Rhizotrogus pallidi-
pennis Blanchard. 
Chrysomelidae 
Crioceris asparagi var. campestris 
Unnaeus (Alcúdia, 20.10.1990). Clava-
reau (1913) showed that Crioceris as-
paragi (Unnaeus), C. campestris Un-
naeus, and C. campestris "Rossius" we-
re synonymous, but on page 41 he ac-
cepted Crioceris macilenta Weise as a 
variety of C. asparagi (L.). White (1981) 
has resolved the taxonomic difficulties 
surrounding C. campestris. The Alcúdia 
specimen was compared with specimens 
in the Natural History Museum, London, 
bearing the names Crioceris asparagi 
P.F. Whitehead, Coleoptera of Mallorca 165 
varo campes tris, and Crioceris macilenta 
Weise. In my view these specimens are 
in reality conspeci1ic, and the correct 
taxonomy 01 the Alcúdia specimen is 
Crioceris macilenta Weise, a species 
already known 1rom the Balearic archi-
pelago (Jolivet, 1953) as well as North 
A1rica, Iberian Peninsula, Sardinia, 
Corsica, Sicily, Italy, Greece (Whitehead, 
in press), and Cyprus. 
Crioceris macilenta Weise shows 
considerable whole-range variation (ove-
rall size, extent 01 elytral vittae, extent 
01 dark pigmentation on appendages 
etc.) which has probably caused past 
misconceptions. 
References 
Clavareau, H. 1913. Coleopterorum 
Catalogus. Pars. 51: Chrysomeli-
dae. W. Junk, Berlin. 103 pp. 
Jolivet, P. 1953. Les Chrysomeloidea 
des ¡les Baleares. Mem. Inst. Roy. 
Sci. Nat. Belgique, 2(50):88. 
Martín-Piera, F. & Coca-Abia, M.M. 
1992. Revisión taxonómica del 
género Rhizotrogus Berthold 1827, 
el grupo de R. cicatricosus Muls. 
1842 (Colo, Melolonthidae). Elytron, 
6:199-220. 
Whitehead, P.F. 1993. Observations on 
Coleoptera 01 Mallorca, Balearic 
Islands. BolI. Soco Hist. Nat. Ba-
lears, 36:45-56. 
Whitehead, P.F. in press. The Coleop-
tera 01 Zakynthos, Greece. 
Hammond, P.M. 1973. Observations on 
British Staphylinidae 3. The British 
species 01 Sepedophilus Gistel 
(Conosomus auctt.). Entomologist's 
mono Mag., 108:130-165. 
White, R.E. 1981. Homonymy in world 
species-group names 01 Criocerinae 
(Coleoptera: Chrysomelidae). U.S. 
Dept. Agric. Tech. Bull. 1629, 1-69. 


Observations on Coleoptera of Mallorca, Balearic Islands (amendments)

Whitehead, P.F.

Revistes Catalanes amb Accés Obert

Observations on Coleoptera 
of Mallorca, Balearic Islands 
(amendments) 
Paul F. WHITEHEAD 
SHNB Whitehead, P.F. 1995. Observations on COleoptera of Mallorca, Balearic Islands (amendments). Bo". Soco Hist. Nat. Balears, 38:163-165. ISSN 0212-260X. Palma de Mallorca. 
~ Diverse ecological and taxonomical aspects of some Coleoptera, which appeared in a previous article, are discussed and commented on. Keywords: Coleoptera, Mallorca, taxomony. 
SOCIET AT D'HISTÓRIA 
NATURAL DE LES BAlEARS 
OBSERVACIONS SOBRE COLEOPTERS DE MALLORCA, ILLES BALEARS 
(PUNTUALlTZACIONS). Es discuteixen i comenten diferents aspectes ecologics 
i taxonomics d'alguns coleopters tractats en un anterior article. 
Paraules e/au: Coleoptera, Mallorca, taxonomia. 
Paul F. WHITEHEAD, Moor Leys, Little Comberton, Pershore, Worcestershire, 
WR10 3EH, England. 
Recepció del manuscrit: 5-set-95; revisió acceptada: 24-oct-95. 
Introduction 
My earlier paper in this Bulletin 
(Whitehead, 1993) discussed a number 
of significant Coleoptera from Mallorca, 
with a strong emphasis on the conser-
vation of species from the crucially 
important north coast wetlands and 
littoral. They include species of Cara-
bidae and Hydrophilidae characteristic of 
productive, regularly inundated fen, an 
extensive prime habitat which contri-
butes to the magnificent varied scenery 
of the area. 
A number of misprints crept into 
my earlier paper through no fault of my 
own, which I am now amending, as well 
as updating knowledge of particular 
species. 
Staphylinidae 
The genus Sepedophilus Gistel. 
Mention was made of two species 
on page 49, both having now been 
studied by Dr. M. Schülke, the leading 
authority of the group. The record of 
Sepedophilus pedicularius (Gravenhorst) 
(Lago Esperanza, Port d'Alcúdia, 
25.10.1990) must be deleted. The spe-
cimen is a female, and according to Dr. 
Schülke cannot therefore be determined, 
but an affinity with Sepedophilus 
164 BolI. Soco Hist. Nat. Balears, 38 (1995) 
. i 
:,: 
.' 
FIg. 1. a) Kemerton, Worcestershire, 
England, 7.8.1992. b) Alcanada, Mallorca, 
Balearic Islands, April 1987. The general 
morphological conformation and microsculptu-
re of the Balearic insect agrees with S. 
testaceus, but turther work on the 
morphology of the genital internal armature 
of a larger sample may, in the author's view, 
provide a basis tor taxonomic revision. 
Fig. 1. a) Kemerton, Worcestershire, Angla-
terra, 7.8.1992. b) Alcanada, Mal/orca, /l/es 
Balears, abril 1987. La conformació morfoló-
gica general així com la microescultura de 
/'insecte balear coincideixen amb els de S. 
testaceus, peró estudis més precisos sobre 
la morfologia de /'armadura genital interna a 
un exemplar més gros podria, segons 
/'autor, donar l/oc a una revisió taxonómica. 
obtusus (Luz e) (Central and southern 
Europe, near East, certainly not Iberian) 
is apparent. 
Dr. Schülke has confirmed my 
identification of Sepedophilus testaceus 
(Fabricius) (coast near lila d'Alcanada, 
April 1987) commenting that the 
structural details of the aedeagus are 
somewhat atypical. Fig. 1 a shows the 
internal structure of an aedeagus of a 
male S. testaceus from Kemerton, 
Worcestershire, England (7.8.1992, at 
Perenniporia fraxinea (Fr.) Ryvarden on 
Fraxinus excelsior Unnaeus), of fairly 
typical appearance (Hammond, 1973) . 
The internal structure of the aedeagus 
of the Alcanada specimen is shown in 
Fig. 1 b. In England, this species is 
characteristic of fungoid angiosperms. 
Its occurrence on Mallorca on Pinus 
halepensis Miller, is more reminiscent of 
Sepedophilus lusitanicus Hammond. 
Pselaphldae 
The entry on page 50 should have 
read as follows: "Brachygluta globuli-
collis aubéii. Tournier. 80 in flood debris 
on beach by Torrent de S'Albufereta, 
21-25.10.1990; Brachygluta schüppelii 
Aubé, a few, Parc de S'Albufera, 
23.10.1990. Examples of both species 
have been seen by Dr. C. Besuchet, 
Geneva". 
Scarabaeldae 
Martín-Piera & Coca-Abia (1992) 
have shown that Rhizotrogus lepidus 
Schaufuss is an insular form of the 
dominantly Iberian Rhizotrogus pallidi-
pennis Blanchard. 
Chrysomelidae 
Crioceris asparagi var. campestris 
Linnaeus (Alcúdia, 20.10.1990). Clava-
reau (1913) showed that Crioceris as-
paragi (Unnaeus), C. campestris Un-
naeus, and C. campestris "Rossius" we-
re synonymous, but on page 41 he ac-
cepted Crioceris macilenta Weise as a 
variety of C. asparagi (L.). White (1981) 
has resolved the taxonomic difficulties 
surrounding C. campestris. The Alcúdia 
specimen was compared with specimens 
in the Natural History Museum, London, 
bearing the names Crioceris asparagi 
P.F. Whitehead, Co/eoptera of Mallorca 165 
varo campes tris, and Crioceris macilenta 
Weise. In my view these specimens are 
in reality conspeci1ic, and the correct 
taxonomy 01 the Alcúdia specimen is 
Crioceris macilenta Weise, a species 
already known 1rom the Balearic archi-
pelago (Jolivet, 1953) as well as North 
A1rica, Iberian Peninsula, Sardinia, 
Corsica, Sicily, Italy, Greece (Whitehead, 
in press), and Cyprus. 
Crioceris macilenta Weise shows 
considerable whole-range variation (ove-
rall size, extent 01 elytral vittae, extent 
01 dark pigmentation on appendages 
etc.) which has probably caused past 
misconceptions. 
References 
Clavareau, H. 1913. Co/eopterorum 
Catalogus. Pars. 51: Chrysomeli-
dae. W. Junk, Berlin. 103 pp. 
Jolivet, P. 1953. Les Chrysomeloidea 
des ¡les Baleares. Mem. Inst. Roy. 
Sci. Nat. Be/gique, 2(50):88. 
Martín-Piera, F. & Coca-Abia, M.M. 
1992. Revisión taxonómica del 
género Rhizotrogus Berthold 1827, 
el grupo de R. cicatricosus Muls. 
1842 (CoL, Melolonthidae). E/ytron, 
6:199-220. 
Whitehead, P.F. 1993. Observations on 
Coleoptera 01 Mallorca, Balearic 
Islands. BolI. Soco Hist. Nat. Ba-
lears, 36:45-56. 
Whitehead, P.F. in press. The Coleop-
tera 01 Zakynthos, Greece. 
Hammond, P.M. 1973. Observations on 
British Staphylinidae 3. The British 
species 01 Sepedophilus Gistel 
(Conosomus auctt.). Entomologist's 
mono Mag., 108:130-165. 
White, R.E. 1981. Homonymy in world 
species-group names 01 Criocerinae 
(Coleoptera: Chrysomelidae). U.S. 
Dept. Agric. Tech. Bull. 1629, 1-69. 


Observations on Coleoptera of Mallorca, Balearic Islands (amendments)

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