The history of life is often viewed as a evenly branching tree; however, in reality it is more like a tangled hedgerow. Many groups of organisms are known to have such a net-like or reticulate evolutionary history, but it is particularly common in ferns and lycophytes (also known as pteridophytes). This dissertation investigates how net-like evolution affects different groups of ferns, with a special emphasis on the model species C-fern (Ceratopteris richardii, also called the antler or water sprite fern). Genomic data are utilized to under-stand hybridization, cryptic species and reticulate evolution in two groups of ferns. The C-fern is shown to be a potential hybrid species, which has important implications for future research using this model organism
