The higher plant NE contains a functional Linker of Nucleoskeleton and Cytoskeleton (LINC) complex based on conserved Sad1-Unc84 (SUN) domain proteins and plant specific Klarsicht/Anc1/Syne homology (KASH) domain proteins (Figure 1). Recent evidence suggests the presence of a plant lamina underneath the inner membrane and various coiled-coil proteins have been hypothesised to be associated with it including Crowded Nuclei (CRWN), Nuclear Envelope Associated Protein (NEAP) protein families as well as the CRWN binding protein KAKU4. In this study, we explore the presence of proteins of these nuclear envelope (NE) proteins from the most ancestral plant species to advanced angiosperms
