Funding This work was supported by the NERC under Grant NE/M010953/1. Acknowledgements Skilled technical support was provided by J. Johnston and J. Bowie. We are very grateful to three reviewers whose comments helped to focus the manuscript. Conceptualization, J.P.; Formal Analysis, J.S., E.H. and J.P.; Investigation, R.M., E.H., J.S. and J.P.; Data Curation, J.S.; Writing, Review & Editing, J.P., E.H. and R.M.; Project Administration, J.P.; Funding Acquisition, J.P. Open access via T&F agreementPeer reviewedPublisher PD