Acknowledgements The authors would like to thank Dr Robin Walker for access to the experimental plots at the SRUC, Craibstone Estate, Aberdeen. JZ was funded by a Natural Environment Research Council (NERC) grant (NE/K016342/1), MOB by a University of Aberdeen Elphinstone Scholarship and by TETFund through Adekunle Ajasin University Akungba (AAUA) Nigeria, YM by a NERC grant (NE/R001529/1) and CGR by a Royal Society University Research Fellowship (URF150571).Peer reviewedPublisher PD
