Funding This work forms part of a NERC New Investigator award for DH (NE/I001743/1), which is gratefully acknowledged. Acknowledgments The authors would like to acknowledge the reviewers, Elizabeth Ritz and Phillip Resor. Their reviews were very constructive, both helping to improve the manuscripts consistency and highlighting a number of errors in the initial submission. The authors would also like to thank Lydia Jagger's keen eye and patience, she helped greatly in removing a number of grammatical errors from the initial draft.Peer reviewedPublisher PD