We introduce an evolutionary metacommunity of multitrophic food webs on
several habitats coupled by migration. In contrast to previous studies that
focus either on evolutionary or on spatial aspects, we include both and
investigate the interplay between them. Locally, the species emerge, interact
and go extinct according to the rules of the well-known evolutionary food web
model proposed by Loeuille and Loreau in 2005. Additionally, species are able
to migrate between the habitats. With random migration, we are able to
reproduce common trends in diversity-dispersal relationships: Regional
diversity decreases with increasing migration rates, whereas local diversity
can increase in case of a low level of dispersal. Moreover, we find that the
total biomasses in the different patches become similar even when species
composition remains different. With adaptive migration, we observe species
compositions that differ considerably between patches and contain species that
are descendant from ancestors on both patches. This result indicates that the
combination of spatial aspects and evolutionary processes affects the structure
of food webs in different ways than each of them alone.Comment: under review at JT