We thank members of the P.V. lab and CBD for helpful discussions and precious help. We thank Alexandre Gehanno, Daan Remans, and Arissa Jokhio for precious help. This work was funded by grants from the European Research Council (NEUROTEMPO), the C1 KU Leuven Internal Funds Program, the EOS Program, the Foundation GENERET, ERANET EPINEURO-DEVO, NSC-Reconstruct, the Belgian FWO, and the Belgian Queen Elizabeth Foundation (P.V.). The authors gratefully acknowledge the VIB Bio Imaging Core for their support and assistance in this work. R.I. was supported by a postdoctoral fellowship of the FRS/FNRS