Bedetti, C

Bellucci, L

Conti, N

Coppola, D

DI CANZIO, E

Pavia, Marco

Petronio, C

Petrucci, M

Salari, L.

Sardella, R

English

Institutional Research Information System University of Turin

Geo.A/p, Val. 2, S. 25-29, 2005 THE LATE PLEISTOCENE VERTEBRATE FAUNA FROM AVETRANA (TARANTO, APULIA, SOUTHERN ITALY): PRELIMINARY REPORT Raffaele Sardella1,2, Claudia Bedettil, Luca Bellucci', Nicoletta Conti', Danilo Coppola3, Emmanuele Di Canzio2, Marco Pavia4 , Carmelo Petronio1,2, Mauro Petrucci' & Leonardo Salari' With 3 figures , Dipartimento di Scienze della Terra, Università di Roma "La Sapienza"; e-mail : raffaele.sardella@uniromal.it 2 CNR-IGAG Istituto di Geologia Ambientale e Geoingegneria 3 Dipartimento di Storia, Università "Tor Vergata" di Roma 4 Dipartimento di Scienze della Terra, Università di Torino Riassunto In una cava inattiva, nei pressi de(la cittadina di Avetrana, a Est di Taranto (Puglia, Italia meridionale), è stata rinvenuta un ricca associazione faunistica a vertebrati contenuta nel riempimento di una cavità di origine carsica. Nella cava affiora la formazione delle "Calcareniti di Gravina", compatta di colore gial­lastro, con ricca malacofauna ed echinidi, ascrivibile al Pleistocene inferiore. A seguito di un saggio di scavo condotto nell'ottobre 2003 è stato possibile condurre una prima analisi stratigrafica del riempi­mento carsico, con campionamento dei diversi livelli riconosciuti e recupero di alcuni degli abbondanti reperti fossili affioranti. Attualmente i resti di vertebrati recuperati sono conservati presso la Soprintendenza di Taranto. Nel riempimento della fessura carsica indagata sono stati riconosciuti nove livelli fossiliferi e due tasche. Le specie presenti nel deposito sono riferibili al Pleistocene superiore. Abstract In an abandoned quarry near Avetrana (Taranto, Apulia, Southern Italy) a fossiliferous karst filling deposit rich in vertebrate remains has been discovered. This deposit fills a wide karst fracture crossing a massive, yel­low marine bio-calcarenite termed "Calcareniti di Gravina" Formation. This formation contains a rich macro­fauna and echinids, referable to the Early Pleistocene. In October 2003, field activities including sampling of the sediments and a preliminary excavation of the fossiliferous levels started . Within the karst filling deposit nine levels and two pockets could be determined. The collected fossils are stored at the "Soprintendenza per i Beni Archeologici per la Puglia" (Taranto, Apulia) . This fossiliferous karst deposit can be referred to the Late Pleistocene. Introduction The Salento Peninsula is well known in the palaeontological literature for its Late Pleistocene vertebrate faunas, mainly in its southern part (Blanc 1920, De Giuli 1983, Corridi 1987, Di Stefano et alii 1992, Bologna et alii 1994, Rustioni et alii 1994 among others). Data available for the lonian northern part of Salento are quite rare. Here we report the discovery of a new fosslliferous locality in the area of Taranto. It is a karst filling deposit rich in fossil bones located in an abandoned cal­carenite quarry, in the area of Avetrana, not far from Manduria (Fig. 1). After a preliminary survey during May 2003, car­ried on by some of the authors (D. Coppola and C. Petronio, in particular), a team of palaeontologists of the "La Sapienza" University, leaded by Prof. 25 I C. Petronio, started the field cam­paign, in accordance with "Soprin­tendenza per i Beni Archeologici per la Puglia". Field work was con­tinued in October 2003, with activities of sampling sediments and a partial excavation of the fossiliferous levels. The collected fossils are stored at "Soprinten­denza per i Beni Archeologici per la Puglia" (Taranto, Apulia). In the present paper we present a preliminary analysis of the col­lected materia I and some generai outlines of the fossiliferous karst deposit. O;....._..;;:5~Okm Stratigraphy The "Calcareniti di Gravina" Formation is a mas­sive bio-calcarenite, rich in molluscs and echinids, that widely outcrops in the central-southern Apulian Peninsula; its age spans the Late Pliocene (Adriatic side) and Early Pleistocene (lonian side) (Ciaranfi et alii 1988). In the considered quarry near Avetrana this formation is exposed in a section which is approximately 10 m thick. The vertebrate fossi I bones occur in a karst fissure filling (Fig. 3). The sediments containing fossil vertebrates are divided into two parts: the upper part fills the main cavity (Iayers 1 to 9), the lower part fills a network of small fissures which opened under the main one. The small fissures (Iayer O in Fig. 3) are filled with orange-yellow nonlaminated sandy clays rich in sm all and medium-sized vertebrate remains. The main cavity is filled with laminated sediments 4,5 to 5,5 m thick. From the bottom to the top the follow­ing levels have been determined (Fig 3): 1) 	 30 cm of clayey sand with rare altered calcare­ous pebbles and bones. A continuous level of calcareous pebbles constitutes the basai part of the layer. 2) 20 cm of sandy clay very rich in fossil remains. 3) 20 cm of clayey sand with some bones and rare calcareous pebbles. 4) 20 cm of sandy clay rich in fossil bones and cal­careous pebbles. This layer is separated from layer 3 by an erosional surface which is marked by a level of calcareous pebbles and bones. Fig. 1: Location of Avetrana 5) 	 140 cm of clayey sand with very abundant bones and rare calcareous pebbles. This layers is char­acterised by a level of bones and pebbles at its base and also by a sandy lens with rare bones 20 cm above t he base of this layer. 6) 	 20 cm of sand with abundant bones and cal­ca reous pebbles. 7) 	 40 cm of clay with abundant bones and large calcareous boulders, especially at the base of the layer (Fig . 2). 8) 	 75 cm of clay and bones, the bones are also con­centrated at the base of the layer, separating it from the underlying layer. 9J 	 70 cm of clay with sparse bones, most of them decalcified. Pa laeontology Most of the fossil remains found at different lev­els of the main cavity are of medium to large size, mainly referable to 805 primigenius (Fig. 3). The pre­liminary analysis of the fossil materia I enables us to present the following faunal list : Layer O: AVES: Perdix perdix, Columba livia, Athene noctua, Pyrrhocorax graculus; MAMMALlA: Erinaceus europaeus, Lepus europaeus, Oryctolagus cuniculus, Hystrix cf. H. cristata, Terricola savi, Felis silvestris. Layer Layer ta, nu Layer lUi ph Layer lUi ph Layer Ce Layer Ce Layer ta, eh Layer Pa Da Layer La'y small sente( mainl lower europ of Te, slight been rabbi t mand reco re 2 to l repres size. In best-r (1-8) ; some chara, and rr Cet and li phala ascrib vores, testifi ers 2 from Geo.A/p, VoI. 2, 2005 26 rana mes lar­~ its i20 cal­lrge the on­9 it lem lev­iize, )re­5 to lene LlA : eus, ata, Layer 1: 80S primigenius. Layer 2: Vulpes vulpes, Canis lupus, Crocuta crocu­ta, Lynx Iynx, Stephanorhinus sp., 80S primige­nius, Bovidae indet., Dama dama, Cervus elaphus. Layer 3: Lepus europaeus, Vulpes vulpes, Canis lupus, 80S primigenius, Dama dama, Cervus ela­phus. Layer 4: Lepus europaeus, Vulpes vulpes, Canis lupus, 80S primigenius, Dama dama, Cervus ela­phus. Layer 5: Vulpes vulpes, Canis lupus, 80S primigenius, Cervus elaphus, Dama dama. Layer 6: Vulpes vulpes, Canis lupus, 80S primigenius, Cervus elaphus, Dama dama. Layer 7: Vulpes vu Ipes, Canis lupus, Crocuta crocu­ta, Stephanorhinus sp., 80S primigenius, Cervus elaphus, Dama dama, ?Megaloceros giganteus. Layer 8: Vulpes vulpes, Canis lupus, Lynx Iynx, Panthera leo, 80S primigenius, Cervus elaphus, Dama dama. Layer 9: fossil bones are absent. Layer ° is characterised by the occurrence of small vertebrate remains. Bird remains are repre­sented by some limb bones, while mammals are mainly represented by teeth . In particular, two lower molar teeth (M 2 and M3) of Erinaceus europaeus, 7 well preserved mandibles and some M, of Terricola sa vi, M' and two of a porcupine, M2 slightly smaller than the living Hystrix cristata, have been recorded. Among lagomorphs the hare and the rabbit occur with some fragments of skull and mandible and some limb bones. Such taxa are also recorded from the mai n cavity deposit (from layers 2 to 8) but are poorly represented. The wild cat is represented by fragmentary limb bones of peculiar size. In the main cavity filling, 80S primigenius is the best-represented taxon in each fossiliferous level (1-8l; layers 5 and 6 are very rich in limb bones, in some cases in anatomical connection. Layer 8 is also characterised by the occurrence of skull fragments and mandibles with jugal teeth. Cervids are represented by some isolated teeth and limb bones (Iayers 2-8); two large-sized first phalanxes recorded in level 7 can probably be ascribed to Megaloceros giganteus. Among carni­vores, the occurrence of the wolf and the red fox is testified by some isolated teeth, occurring from lay­ers 2 to 8, the Iynx and the cave lion are recorded from layer 8 (some teeth and a talus respectively). Fig . 2: Fossil bones of layer 7 (scale bar: 20 cm). The occurrence of a rhino is testified in layers 2 and 7. This taxon is represented by one fragmentary molar tooth and one pisiform in every layer. The faunal assemblage on the whole can be referred to the Late Pleistocene. The occurrence of the fallow deer (in particular of the modern sub­species Dama dama dama) and of a rhino, generally referable to Stephanorhinus sp., recorded from lay­ers 2 and 7, gives important biochronological con­straints. In fact, the modern fallow deer was wide­spread in Italy at the beginning of the Late Pleistocene, while rhinos referable to the genus Stephanorhinus survived until the beginning of the Pleniglacial (MIS 3) (Gliozzi et alii 1997). At the moment, only generai considerations on the palaeoenvironmental conditions can be pointed out. In layer 0, the occurrence of Terricola sa vii and Hystrix cf. H. cristata suggests the presence of tem­perate climatic conditions with dry and open palaeoenvironments. Moreover, such a generai framework is supported also by the occurrence of the avifauna including Perdix perdix and Athene noctua, while Columba livia and Pyrrhocorax graculus sug­gest the presence of rocky cliffs. In the sequence fill­ing the main cavity (layer 1 to 8), large mammal species of a wider ecological significance occur. Preliminary taphonomical observations indicate that the fossil bones seem to be not oriented. This n05 Geo.A/p, VoI. 2, 2005 27 l 9 8 7 6 I~:~~~:'J Cloys I~;~~~{~I Sandy clays 5 1"'1Sand. liit;'~l Cloyed SCII1ds D fossi I bone.s 4 Io ~I Colcore.ous pe.bb les 3 2 D Strat ified deposi t lZ1 Not stratified depos it Fig . 3: Stratigraphy of the fossiliferous deposit fact suggests quick deposition of the fossil-bearing sediments, which is also supported by sedimento­logical observations. In fact, the different layers are characterised by a normally graded distribution of the sediment, with the heaviest material like big bones and calcareous pebbles concentrated in the lowest part of the layer. Frequently such bones and pebbles constitute a well defined level at the base of the layer to separate one layer from the underly­ing one. A detailed analysis of the fossi I remains has just begun, in accordance with the "Soprintendenza ai Beni Archeologici per la Puglia", with the aim of providing further palaeontological information and a framework of the palaeoenvironmental evolution of the area during the Late Pleistocene. Acknowledgments Ali phases of fieldwork were financially support­ed by CNR - IGAG and by the Municipality of Avetrana . A special mention is due to Francesco Nigro, responsible for the Cultural Heritage. The support of Maria Antonietta Gorgoglione, responsi­ble for "Soprintendenza per i Beni Archeologici per la Puglia", is warmly acknowledged. Moreover, the Earth Science Department of the University of Turin and the "Museo delle Civiltà Preclassiche delle Murge Meridionali" provided facilities for research. We wish to thank Giuseppe "Pippo" Arcidiacono, Francesco Ciminelli, Vincenza Montenegro and Michael Giagnoni for participating the field work and, finally, Karl Krainer and Marzia Breda for their comments and suggestions on the manuscript. References Blanc, G. A.(1920) : Grotta Romanelli . - Arch. Antrop. Etn., 50( 1-4): 1-39. Bologna, P., Di Stefano, G., Manzi, G., Petronio, c., Sardella, R., Squazzini, E. (1994) : Late Pleistocene mammals from the Melpignano (Le) "Ventarole": pre­liminary analysis and correlations. - BolI. Soc. Paleont. It., 33 (2): 265-274. Geo.Alp, VoI. 2, 2005 Ciarar Ge tro Corrid faL Mu De Gil La COI Di St( Sql oss (Lei Gliozz Car G., 28 I.. sit 1depoSi t geo The :sponsi­gici per ver, the Df Turin e delle ~search. liacono, ro and Id work or their pt. 'op. Etn., mio, c., istocene le": pre­Paleont. . 2,2005 Ciaranfi, N., Pieri, P., Ricchetti, G. (1988): Note alla Carta Geologica delle Murge e del Salento (Puglia cen­tromeridionale). - Mem. Soc. Geol. It., 41 (1): 449-460. Corridi, C. (1987): Faune pleistoceniche del Salento: 2. La fauna di fondo Cattìe, Maglie, Lecce. - Quaderni del Museo Comunale di Paleontologia, 3: 5-74. De Giuli, C. (1983): Le faune pleistoceniche del Salento: 1. La fauna di S. Sidero 3. - Quaderni del Museo Comunale di Paleontologia, 1: 45-84. Di 	 Stefano, G., Petronio, C., Sardella, R., Savelloni, V., Squazzini, E. (1992): Nuove segnalazioni di brecce ossifere nella costa fra Castro Marina e Otranto (Lecce). - Il Quaternario, 5 (1): 3-10. Gliozzi, E., Abbazzi, L., Argenti, P., Azzaroli, A., Caloi, L., Ca passo Barbato, L., Di Stefano, G., Esu, D., Ficcarelli, G., Girotti, O., Kotsakis, l, Masini, F., Mazza, P., Mezzabotta C., Palombo, M. R., Petronio, c., Rook, L., Sala, B., Sardella, R., Zanalda, E., Torre, D. (1997): Biochronology of selected Mammals, Molluscs and Ostracods from the Middle Pliocene to the Late Pleistocene in Italy. The state of the art. - Riv. Ital. Paleont. Strat., 103(3): 369-388. Rustioni, M., Mazza, P., Abbazzi, L., Delfino, M., Rook, L., Petrucci, S., Vianello, F. (1994) : The Wurmian fauna from Sternatia (Lecce, Apulia, Italy). - Boli. Soc. Paleont. It., 33 (2): 279-288. Manuscript submitted: December 17, 2004 Revised manuscript accepted: February 22, 2005 Geo.A/p, VoI. 2, 2005 29 

The Late Pleistocene vertebrate fauna from Avetrana (Taranto, Apulia, Southern Italy): Preliminary report.

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The Late Pleistocene vertebrate fauna from Avetrana (Taranto, Apulia, Southern Italy): Preliminary report.

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