Stems and Spectral Sequences

We introduce the category Pstem[n] of n-stems, with a functor P[n] from spaces to Pstem[n]. This can be thought of as the n-th order homotopy groups of a space. We show how to associate to each simplicial n-stem Q an (n+1)-truncated spectral sequence. Moreover, if Q=P[n]X is the Postnikov n-stem of a simplicial space X, the truncated spectral sequence for Q is the truncation of the usual homotopy spectral sequence of X. Similar results are also proven for cosimplicial n-stems. They are helpful for computations, since n-stems in low degrees have good algebraic models

Inherited biotic protection in a Neotropical pioneer plant

Chelonanthus alatus is a bat-pollinated, pioneer Gentianaceae that clusters in patches where still-standing, dried-out stems are interspersed among live individuals. Flowers bear circum-floral nectaries (CFNs) that are attractive to ants, and seed dispersal is both barochorous and anemochorous. Although, in this study, live individuals never sheltered ant colonies, dried-out hollow stems - that can remain standing for 2 years - did. Workers from species nesting in dried-out stems as well as from ground-nesting species exploited the CFNs of live C. alatus individuals in the same patches during the daytime, but were absent at night (when bat pollination occurs) on 60.5% of the plants. By visiting the CFNs, the ants indirectly protect the flowers - but not the plant foliage - from herbivorous insects. We show that this protection is provided mostly by species nesting in dried-out stems, predominantly Pseudomyrmex gracilis. That dried-out stems remain standing for years and are regularly replaced results in an opportunistic, but stable association where colonies are sheltered by one generation of dead C. alatus while the live individuals nearby, belonging to the next generation, provide them with nectar; in turn, the ants protect their flowers from herbivores. We suggest that the investment in wood by C. alatus individuals permitting stillstanding, dried-out stems to shelter ant colonies constitutes an extended phenotype because foraging workers protect the flowers of live individuals in the same patch. Also, through this process these dried-out stems indirectly favor the reproduction (and so the fitness) of the next generation including both their own offspring and that of their siblings, alladding up to a potential case of inclusive fitness in plants

Growth allocation and stand structure in Norway spruce stands

This thesis is based on analyses of permanent sample plot data gathered over periods of 10-34 years from an experiment in which a wide range of thinning regimes, and thinnings combined with N-fertilisation, were applied to 25 even-aged Norway spruce (Picea abies (L.) Karst.) stands in southern Sweden (56-63 No). At the start of the experiment, before the first thinning, the dominant height was 12-18 m. The overall objective was to evaluate the extent to which growth allocation along the bole and the stand structure of Picea abies stands can be controlled by different silvicultural regimes. To do this the data were used in four studies to evaluate the impact of: thinning and N-fertilisation on stem form and taper (Study I); different thinning regimes on the removal and growth in the diameter at breast height (DBH) of individual stems (Studies II and III); and the thinning regimes on the growth in mean DBH of four classes of the largest stems by DBH ha 1 (Study IV). The studies (ii) and (iii) form a growth model. In stands subjected to different thinning regimes, one model predicts which individual trees will remain at future points in time and an associated model predicts the future DBH of the remaining stems. Separate models were developed for stands thinned from below, stands thinned from above and unthinned stands. In Study IV the actual and genuine increases in the arithmetic mean DBH of the 100, 200, 300 and 400 largest stems by DBH ha 1 associated with six different thinning regimes in periods up to 35 years were compared to the corresponding stems in unthinned stands. The goals of achieving rapid diameter growth and low stem tapering cannot be attained simultaneously as heavy thinnings cause increased tapering, and thus silvicultural regimes must reflect a compromise between these and other production objectives. Trees in thinned and N-fertilised stands had the same taper as trees in equally thinned, unfertilised stands. Heavy thinnings from below promote high frequencies of thick stems and extra heavy thinnings promote high frequencies of extra thick stems. Thinning from above (or no thinnngs) may be an alternative to thinning from below in situations where a main crop consisting of moderately thick stems would be regarded as a satisfactory outcome. The actual mean DBH of larger stems can be increased, compared to the corresponding stems in unthinned stands, by up to 2.6 mm per year if extra heavy thinnings are carried out. The biological response to thinning of thick stems is influenced by the thinning intensity but not by the thinning method. The variation in DBH increases over time but increases more in stands thinned from above and unthinned stands than in stands thinned from below

Hungarian neutral vowels

In Hungarian, stems containing only front unrounded (neutral) vowels fall into two groups: one group taking front suffixes, the other taking back suffixes in vowel harmony. The distinction is traditionally thought of as purely lexical. Beňuš and Gafos (2007) have recently challenged this position, claiming that there are significant articulatory differences between the vowels in the two groups. Neutral vowels also occur in vacillating stems. These typically contain one back vowel and one or more neutral vowels, and accept both front and back suffixes, with extensive inter- and intra-speaker variation. Based on Beňuš and Gafos’s line of argument, the expectation is that vacillating stems will display a kind of phonetic realisation that is distinct from both harmonic and anti-harmonic stems. We present the results of an ongoing acoustic study on the acoustics of neutral vowels, partly re-creating Beňuš and Gafos’s conditions, but also including vacillating stems. To map the extent of individual and dialectal variation regarding vacillating stems, a grammaticality judgement test was also carried out on speakers of two dialects of Hungarian, crucially differing in the surface inventory of neutral vowels. We present our first findings about how this phonetic difference influences the phonological behaviour of vacillating stems

Holger Pedersen's "Études lituaniennes" revisited

Holger Pedersen’s "Études lituaniennes" reflects the issues under discussion at the time of its publication (1933). Its five unequal chapters deal with the following topics: I. The Lithuanian future and its Indo-European origins: the sigmatic formation, the 3rd person zero ending, the short root vowels e and a, the shortening and metatony in the 3rd person, and the future participle. II. The accentuation of nouns in Lithuanian: accentual mobility in the Indo- European consonant stems and its absence in the o-stems, the origins of accentual mobility in Lithuanian nominal paradigms, the accentuation of separate case forms, and accentual peculiarities of the adjective. III. The acute tone of the root in consonant stems. IV. The past active participle. V. Secondary vocalic alternations: new vowel length and new acute tone

First report of Desmiphora hirticollis (Olivier) (Coleoptera: Cerambycidae) on Wigandia urens (Ruiz and Pavón) H.B.K. (Hydrophilaceae) in Mexico

Desmiphora hirticollis (Olivier) (Coleoptera: Cerambycidae) was found in Oaxaca, Mexico, feeding in the stems of Wigandia urens (Ruiz and Pavón) H.B.K. (Hydrophilaceae), a new host record. Information about damage in the plant stems and some observations of the insect’s biology are provided

Temperature and inoculation method influence disease phenotypes and mortality of Eucalyptus marginata clonal lines inoculated with Phytophthora cinnamomi

Survival of 1-year-old plants of three clonal lines of Eucalyptus marginata (jarrah), two ranked as resistant (RR1 and RR2) and one as susceptible (SS1) to Phytophthora cinnamomi, was assessed after pathogen inoculation with either mycelial mats underbark or zoospores on the stem. Plants were grown at 15, 20, 25 and 30°C. Method of inoculation did not produce comparable mortalities of the clonal lines, particularly at 25 and 30°C. At these temperatures, all three clonal lines had 100% mortality when inoculated underbark, but when inoculated with zoospores, RR1 had 60% survival and lines SS1 and RR2 had 100% mortality. Generally, the level of resistance of all clonal lines declined with increasing temperature. RR2 had consistently higher mortality than SS1, and is therefore not considered resistant. Lesion development was also measured in detached stems of RR1 and a susceptible clonal line (SS2) each inoculated underbark with four different P. cinnamomi isolates. Stems were assessed for lesion development at 20, 25 and 30°C for 4 days. For all four isolates, detached stems of RR1 generally had smaller lesions than those of SS2, particularly at 30°C. The increase in lesion length with increasing temperature was greatest for SS2. Detached stems may have potential in screening for jarrah resistant toP. cinnamomi and allow identification of susceptible clonal lines at 30°C

Impact of \u3ci\u3eMecinus Janthinus\u3c/i\u3e (Coleoptera: Curculionidae) on the Growth and Reproduction of \u3ci\u3eLinaria Dalmatica\u3c/i\u3e (Scrophulariaceae)

Dalmatian toadflax, Linaria dalmatica (L.) Mill. (Scrophulariaceae), a native to the eastern Mediterranean and Black Sea regions of Europe and Asia, has invaded over one million hectares in the western United States and Canada, in habitats similar to its native range. Once established, the aggressive vegetative growth of the plant allows it to invade undisturbed habitats where it can out-compete most other vegetation, placing native plant communities at risk. Biological control of L. dalmatica with Mecinus janthinus Thomson (Coleoptera: Curculionidae) has shown promise in the field. In both studies reported in this paper, the presence of insect attack reduced L. dalmatica plant growth and reduced plant reproductive potential. In a field sleeve cage study, insect- attacked stems were significantly shorter (18 cm) and had 50-70% fewer fruits and flowers than the control stems at the end of the study period. M. janthinus attacked stems showed little apical growth, fewer fruits and flowers, and lower stem biomass relative to control stems. Similar results were observed in the potted plant study where the influence of the extensive root system of the plant was eliminated. This negative impact by the insect is caused both by adult feeding in the apical portion of the plant and the physical destruction of the plant stem from larvae feeding. The decrease in the insect-attacked stem heights may also have an impact on seed dispersal from the mature reproductive structures. A combination of decreased seed production through M. janthinus biological control and poor seedling competition in the moisture limited sites common to north-central Washington State and other similarly dry habitats may negatively influence L. dalmatica populations more than general models predict