Rodent Dental Microwear Texture Analysis as a Proxy for Fine-Scale Paleoenvironment Reconstruction

Dental microwear texture analysis (DMTA) of fossil fauna has become a valuable tool for dietary inference and paleoenvironment reconstruction. Most of this work has utilized larger taxa with larger home ranges. These studies may result in broader-scale habitat inferences that could mask the details of complex mosaic habitats. Rodent DMTA offers an opportunity to work at finer spatial scales because most species have smaller home ranges. Rodents are also keystone species within their ecosystems, abundant, ubiquitous, and found in many fossil deposits. These attributes make them excellent proxies for environmental reconstructions. However, the application of DMTA to rodents remains relatively new. Furthermore, many rodent species are dietary generalists, and individuals available for study in museum collections lack detail on feeding behavior, which makes it difficult to develop strong dietary associations with microwear patterns. The same holds for limited environmental metadata associated with such samples. This dissertation sought to explore the efficacy of rodent DMTA as a proxy for fine-scale paleoenvironment reconstruction and to establish a baseline of extant incisor and molar textures with detailed metadata associations to aid in future comparisons to fossil taxa. The biomonitoring project at Kolomela Mine, located within South Africa’s Northern Cape, provided an ideal opportunity with which to conduct this research. Stomach content analyses conducted on 214 muroid specimens caught within the Kolomela properties examined diets by species, location, and month. These analyses indicated that the Kolomela rodent community mainly consumed grass seed despite the presence of other foodstuff within stomachs. A confocal profiler scanned high resolution casts to provide microwear textures for 198 incisors and 175 molars, from which SSFA and ISO parameter data were derived. Statistical tests explored the effects of diet, taxon, and habitat attributes on the central tendencies of these parameters, as well as effects by tooth form. Incisor microwear textures seemed to possess a stronger environmental signal than that of molars, with analyses indicating significant variation by species, macrohabitat, microhabitat, burrowing behavior, soil, and land cover classification. These results suggested that while soil characteristics had a strong influence on parameter central tendencies, incisor microwear textures seem to result from complex interactions with habitat characteristics. Molar microwear did not parse the considered dietary categories, likely because all individuals had diets dominated by grass seed that swamped any diet signal reflecting the food elements of each group. Significant variation in parameter central tendencies by both species and burrowing behavior were believed to be the result of differing molar topography between Gerbillinae and Murinae specimens. Analyses also separated molar microwear from different dust levels, which indicated that perhaps an environmental signal can be parsed, at least when diets are homogeneous and controlled for. Finally, molar and incisor microwear textures were significantly different from one another, presumably due to a) different roles in food acquisition and process, b) different rates in gross wear and surface turnover, and c) different degrees of interaction with exogenous grit and the outside environment. These results suggest that both tooth types should be considered in future paleoenvironment reconstructions

ALMA Observations of a Candidate Molecular Outflow in an Obscured Quasar

We present Atacama Large Millimeter/Submillimeter Array (ALMA) CO (1-0) and CO (3-2) observations of SDSS J135646.10+102609.0, an obscured quasar and ultra-luminous infrared galaxy (ULIRG) with two merging nuclei and a known 20-kpc-scale ionized outflow. The total molecular gas mass is M_{mol} ~ 9^{+19}_{-6} x 10^8 Msun, mostly distributed in a compact rotating disk at the primary nucleus (M_{mol} ~ 3 x 10^8 Msun) and an extended tidal arm (M_{mol} ~ 5 x 10^8 Msun). The tidal arm is one of the most massive molecular tidal features known; we suggest that it is due to the lower chance of shock dissociation in this elliptical/disk galaxy merger. In the spatially resolved CO (3-2) data, we find a compact (r ~ 0.3 kpc) high velocity (v ~ 500 km/s) red-shifted feature in addition to the rotation at the N nucleus. We propose a molecular outflow as the most likely explanation for the high velocity gas. The outflowing mass of M_{mol} ~ 7 x 10^7 Msun and the short dynamical time of t_{dyn} ~ 0.6 Myr yield a very high outflow rate of \dot{M}_{mol} ~ 350 Msun/yr and can deplete the gas in a million years. We find a low star formation rate (< 16 Msun/yr from the molecular content and < 21 Msun/yr from the far-infrared spectral energy distribution decomposition) that is inadequate to supply the kinetic luminosity of the outflow (\dot{E} ~ 3 x 10^43 erg/s). Therefore, the active galactic nucleus, with a bolometric luminosity of 10^46 erg/s, likely powers the outflow. The momentum boost rate of the outflow (\dot{p}/(Lbol/c) ~ 3) is lower than typical molecular outflows associated with AGN, which may be related to its compactness. The molecular and ionized outflows are likely two distinct bursts induced by episodic AGN activity that varies on a time scale of 10^7 yr.Comment: 16 pages, 7 figures, ApJ accepte

Estimating Black Hole Masses in Active Galaxies Using the Halpha Emission Line

It has been established that virial masses for black holes in low-redshift active galaxies can be estimated from measurements of the optical continuum strength and the width of the broad Hbeta line. Under various circumstances, however, both of these quantities can be challenging to measure or can be subject to large systematic uncertainties. To mitigate these difficulties, we present a new method for estimating black hole masses. From analysis of a new sample of broad-line active galactic nuclei, we find that Halpha luminosity scales almost linearly with optical continuum luminosity and that a strong correlation exists between Halpha and Hbeta line widths. These two empirical correlations allow us to translate the standard virial mass system to a new one based solely on observations of the broad Halpha emission line.Comment: to appear in Apj; 8 pages; 5 figures; uses emulateapj5.st

Observations of Feedback from Radio-Quiet Quasars: I. Extents and Morphologies of Ionized Gas Nebulae

Black hole feedback -- the strong interaction between the energy output of supermassive black holes and their surrounding environments -- is routinely invoked to explain the absence of overly luminous galaxies, the black hole vs. bulge correlations and the similarity of black hole accretion and star formation histories. Yet direct probes of this process in action are scarce and limited to small samples of active nuclei. We present Gemini IFU observations of the distribution of ionized gas around luminous, obscured, radio-quiet (RQ) quasars at z~0.5. We detect extended ionized gas nebulae via [O III]5007 emission in every case, with a mean diameter of 28 kpc. These nebulae are nearly perfectly round. The regular morphologies of nebulae around RQ quasars are in striking contrast with lumpy or elongated nebulae seen around radio galaxies at low and high redshifts. We present the uniformly measured size-luminosity relationship of [O III] nebulae around Seyfert 2 galaxies and type 2 quasars spanning 6 orders of magnitude in luminosity and confirm the flat slope of the correlation (R ~ L^{0.25+/-0.02}). We find a universal behavior of the [O III]/H-beta ratio in our entire RQ quasar sample: it persists at a constant value (~10) in the central regions, until reaching a "break" isophotal radius ranging from 4 to 11 kpc where it starts to decrease. We propose a model of clumpy nebulae in which clouds that produce line emission transition from being ionization-bounded at small distances from the quasar to being matter-bounded in the outer parts of the nebula, which qualitatively explains the observed line ratio and surface brightness profiles. It is striking that we see such smooth and round large-scale gas nebulosities in this sample, which are inconsistent with illuminated merger debris and which we suggest may be the signature of accretion energy from the nucleus reaching gas at large scales.Comment: 44 pages, 11 figures, 3 tables. Accepted for publication in MNRA

Observations of Feedback from Radio-Quiet Quasars - II. Kinematics of Ionized Gas Nebulae

The prevalence and energetics of quasar feedback is a major unresolved problem in galaxy formation theory. In this paper, we present Gemini Integral Field Unit observations of ionized gas around eleven luminous, obscured, radio-quiet quasars at z~0.5 out to ~15 kpc from the quasar; specifically, we measure the kinematics and morphology of [O III]5007 emission. The round morphologies of the nebulae and the large line-of-sight velocity widths (with velocities containing 80% of the emission as high as 1000 km/s combined with relatively small velocity difference across them (from 90 to 520 km/s) point toward wide-angle quasi-spherical outflows. We use the observed velocity widths to estimate a median outflow velocity of 760 km/s, similar to or above the escape velocities from the host galaxies. The line-of-sight velocity dispersion declines slightly toward outer parts of the nebulae (by 3% per kpc on average). The majority of nebulae show blueshifted excesses in their line profiles across most of their extents, signifying gas outflows. For the median outflow velocity, we find a kinetic energy flow between 4x10^{44} and 3x10^{45} erg/s and mass outflow rate between 2000 and 20000 Msun/yr. These values are large enough for the observed quasar winds to have a significant impact on their host galaxies. The median rate of converting bolometric luminosity to kinetic energy of ionized gas clouds is ~2%. We report four new candidates for "super-bubbles" -- outflows that may have broken out of the denser regions of the host galaxy.Comment: 23 pages, 10 figures, 2 tables, accepted for publication in MNRA

Dynamin: Membrane Scission Meets Physics

SummaryDynamin hydrolyzes GTP to constrict and sever membranes. Experimental advances bring dynamin into the realm of physics and reveal key roles for membrane tension and bending at the edge of the constriction

SDSS-IV MaNGA: Identification of active galactic nuclei in optical integral field unit surveys

In this paper, we investigate 2727 galaxies observed by MaNGA as of June 2016 to develop spatially resolved techniques for identifying signatures of active galactic nuclei (AGN). We identify 303 AGN candidates. The additional spatial dimension imposes challenges in identifying AGN due to contamination from diffuse ionized gas, extra-planar gas and photoionization by hot stars. We show that the combination of spatially-resolved line diagnostic diagrams and additional cuts on H$\alpha$ surface brighness and H$\alpha$ equivalent width can distinguish between AGN-like signatures and high-metallicity galaxies with LINER-like spectra. Low mass galaxies with high specific star formation rates are particularly difficult to diagnose and routinely show diagnostic line ratios outside of the standard star-formation locus. We develop a new diagnostic -- the distance from the standard diagnostic line in the line-ratios space -- to evaluate the significance of the deviation from the star-formation locus. We find 173 galaxies that would not have been selected as AGN candidates based on single-fibre spectral measurements but exhibit photoionization signatures suggestive of AGN activity in the MaNGA resolved observations, underscoring the power of large integral field unit (IFU) surveys. A complete census of these new AGN candidates is necessary to understand their nature and probe the complex co-evolution of supermassive black holes and their hosts.Comment: 18 pages, 11 figures, accepted to MNRA

A New Sample of Low-mass Black Holes in Active Galaxies

We present an expanded sample of low-mass black holes (BHs) found in galactic nuclei. Using standard virial mass techniques to estimate BH masses, we select from the Fourth Data Release of the Sloan Digital Sky Survey all broad-line active galaxies with masses < 2 x 10^6 M_sun. BHs in this mass regime provide unique tests of the relationship between BHs and galaxies, since their late-type galaxy hosts do not necessarily contain classical bulges. Furthermore, they provide observational analogs of primordial seed BHs and are expected, when merging, to provide strong gravitational signals for future detectors such as LISA. From our preliminary sample of 19, we have increased the total sample by an order of magnitude to 174, as well as an additional 55 (less secure) candidates. The sample has a median BH mass of = 1.3 x 10^6 M_sun, and in general the objects are radiating at high fractions of their Eddington limits. We investigate the broad spectral properties of the sample; 55 are detected by \rosat, with soft X-ray luminosities in the range 10^40 to 7 x 10^43 ergs/sec. Much like the preliminary sample, these objects are predominantly radio-quiet (R = f_6cm/f_4400A < 10), but 11 objects are detected at 20 cm, with radio powers (10^21-10^23 W/Hz) that may arise from either star formation or nuclear activity; only 1% of the sample is radio-loud. We further confirm that, with =-19.3 and = 0.7 mag, the host galaxies are low-mass, late-type systems. At least 40% show disk-like morphologies, and the combination of host galaxy colors and higher-order Balmer absorption lines indicate intermediate-age stellar populations in a subset of the sample.Comment: to appear in ApJ; 13 pages, 8 figure

The Host Galaxy and Central Engine of the Dwarf AGN POX 52

We present new multi-wavelength observations of the dwarf Seyfert 1 galaxy POX 52 in order to investigate the properties of the host galaxy and the active nucleus, and to examine the mass of its black hole, previously estimated to be ~ 10^5 M_sun. Hubble Space Telescope ACS/HRC images show that the host galaxy has a dwarf elliptical morphology (M_I = -18.4 mag, Sersic index n = 4.3) with no detected disk component or spiral structure, confirming previous results from ground-based imaging. X-ray observations from both Chandra and XMM show strong (factor of 2) variability over timescales as short as 500 s, as well as a dramatic decrease in the absorbing column density over a 9 month period. We attribute this change to a partial covering absorber, with a 94% covering fraction and N_H = 58^{+8.4}_{-9.2} * 10^21 cm^-2, that moved out of the line of sight in between the XMM and Chandra observations. Combining these data with observations from the VLA, Spitzer, and archival data from 2MASS and GALEX, we examine the spectral energy distribution (SED) of the active nucleus. Its shape is broadly similar to typical radio-quiet quasar SEDs, despite the very low bolometric luminosity of L_bol = 1.3 * 10^43 ergs/s. Finally, we compare black hole mass estimators including methods based on X-ray variability, and optical scaling relations using the broad H-beta line width and AGN continuum luminosity, finding a range of black hole mass from all methods to be M_bh = (2.2-4.2) * 10^5 M_sun, with an Eddington ratio of L_bol/L_edd = 0.2-0.5.Comment: 19 pages, 16 figures, accepted for publication in Ap