Composition of canopy ants (Hymenoptera: Formicidae) at Ton Nga Chang Wildlife Sanctuary, Songkhla Province, Thailand

) on each study site. Pyrethroid fogging was two monthly applied to collect ants on three trees at random in a permanent plot. A total of 118 morphospecies in 29 genera belonging to six subfamilies were identified. The Formicinae subfamily found the highest species numbers (64 species) followed by Myrmicinae (32 species), Pseudomyrmecinae (10 species), Ponerinae (6 species), Dolichoderinae (5 species) and Aenictinae (1 species). Myrmicinae and Ponerinae showed a significant difference of mean species number between sites (P<0.05) while Formicinae and Myrmicinae also showed a significant difference of mean species number between months (P<0.05). However, there were no interactions between sites and months in any subfamily

Aenictus sulawesiensis Jaitrong & Wiwatwitaya 2013, new species

Aenictus sulawesiensis, new species (Fig. 2 A–C) Material examined. — Holotype worker from Indonesia, S. Sulawesi, Barru, Taneterilau, Lipukasi, Forest Complex Coppo (4°30'S, 119°37'E), 8 Jan.2011, coll. Sk. Yamane, CE11-SKY-21 (MZB). Sixty-seven paratype workers, same data as holotype (AMK, BMNH, MCZC, MHNG, MZB, SKYC, THNHM). Measurements. — Holotype : TL 3.25 mm; HL 0.76 mm; HW 0.65 mm; SL 0.63 mm; ML 1.05 mm; PL 0.29 mm, CI 85; SI 96. Paratypes (n = 9): TL 3.25–3.30 mm; HL 0.75–0.78 mm; HW 0.63–0.65 mm; SL 0.60–0.63 mm; ML 1.04–1.06 mm; PL 0.28–0.30 mm, CI 82–85; SI 96–98. Worker description. — Head in full-face view elliptical, distinctly longer than broad, with feebly convex sides; posterior margin convex; occipital margin bearing a narrow collar. Antennal scape relatively short, extending beyond 2/3 of head length, but not reaching posterolateral corner of head; antennal segment II slightly shorter than broad; III–VII each almost as long as broad; terminal segment almost as long as VII+VIII+IX. Frontal carinae well developed, fused at the level of antennal base to form a single carina, extending less than half length of head; posterior half of frontal carina very poorly developed, with head in profile roundly concave. Parafrontal ridge well developed, reaching 1/3 of head length (0.30 mm); seen in profile, its anteriormost part well developed and subtriangular, and posterior part feebly convex. Masticatory margin of mandible with large apical tooth followed by a medium-sized subapical tooth and 5–6 denticles; basal margin lacking denticles. Mesosoma elongate and stout; promesonotum seen in profile slightly convex dorsally, sloping gradually to metanotal groove; mesopleuron not clearly demarcated from metapleuron; metanotal groove indistinct; propodeum in profile lower than promesonotum, nearly straight dorsally; propodeal junction angulate; declivity of propodeum shallowly concave, encircled with a rim. Petiole subsessile, almost as long as high, its node short and elevated posteriorly; subpetiolar process weakly developed or almost absent, its ventral margin feebly convex. Postpetiole almost as long as petiole, dorsally convex. Dorsum of head punctate; lateral face with weaker punctation (reticulate with smooth and shiny bottoms) than dorsum and partly smooth and shiny or superficially reticulate with smooth interspaces. Antennal scape microreticulate. Mandible entirely micropunctate except for apical tooth and along masticatory margin. Greater part of pronotum superficially sculptured or smooth and shiny. Petiole entirely punctate and opaque; postpetiole entirely punctate except small area on dorsum shiny. First gastral tergite and sternite smooth and shiny except for the basalmost part with dense punctures. Basal half of femora microreticulate, but apical half superficially macroreticulate, smooth and shiny, partly superficially shagreened with smooth and shiny interspaces. Tibiae microreticulate, somewhat shiny. Head and mesosoma dorsally with dense standing hairs; longest pronotal hair 0.23–0.25 mm long. Dorsum of head, mandible and mesosoma dark brown; legs, waist, and gaster dark reddish brown to reddish brown; antennal scape dark brown except for apicalmost portion reddish brown; all funicular segments reddish brown. Typhlatta spot absent. Etymology. — The specific name is derived from name of the type locality, Sulawesi Island of Indonesia. Distribution. — Sulawesi (Fig. 3). Notes. — So far A. sulawesiensis has been known only from the type locality. This species is very similar to A. kutai (see under A. kutai).Published as part of Jaitrong, Weeyawat & Wiwatwitaya, Decha, 2013, Two New Species Of The Aenictus Pachycerus Species Group (Hymenoptera: Formicidae: Aenictinae) From Southeast Asia, pp. 97-102 in Raffles Bulletin of Zoology 61 (1) on pages 100-101, DOI: 10.5281/zenodo.450930

タイ ニ オケル ドジョウ セッソク ドウブツ グンシュウ エノ シンリン カクラン ノ コウカ

京都大学0048新制・論文博士博士(農学)乙第11677号論農博第2560号新制||農||913(附属図書館)学位論文||H17||N4060(農学部図書室)23490UT51-2005-D595(主査)教授 武田 博清, 教授 高藤 晃雄, 教授 東 順一学位規則第4条第2項該当Doctor of Agricultural ScienceKyoto UniversityDA

Aenictus carolianus Zettel & Sorger 2010

Aenictus carolianus Zettel & Sorger, 2010: 4 paratype workers from the Philippines, Cebu City, Cantipla-I Forest Reserve, 1 Mar.2008, coll. H. Zettel & C. V. Pangantihon, #512 (SKYC and THNHM).Published as part of Jaitrong, Weeyawat & Wiwatwitaya, Decha, 2013, Two New Species Of The Aenictus Pachycerus Species Group (Hymenoptera: Formicidae: Aenictinae) From Southeast Asia, pp. 97-102 in Raffles Bulletin of Zoology 61 (1) on page 101, DOI: 10.5281/zenodo.450930

Aenictus powersi Wheeler & Chapman 1930

Aenictus powersi Wheeler & Chapman in Wheeler, 1930: 11 syntype workers (four pins, two on a pin, three on each of remaining pins) from the Philippines, Negros, Dumaguete, 540 m (MCZC).Published as part of Jaitrong, Weeyawat & Wiwatwitaya, Decha, 2013, Two New Species Of The Aenictus Pachycerus Species Group (Hymenoptera: Formicidae: Aenictinae) From Southeast Asia, pp. 97-102 in Raffles Bulletin of Zoology 61 (1) on page 101, DOI: 10.5281/zenodo.450930

Aenictus reyesi Chapman 1963

<p> <i>Aenictus reyesi</i> Chapman, 1963:</p> <p>20 snytypes from the Philippines, Negros, Horns of Negros, 450 m (MCZC).</p>Published as part of <i>Jaitrong, Weeyawat & Wiwatwitaya, Decha, 2013, Two New Species Of The Aenictus Pachycerus Species Group (Hymenoptera: Formicidae: Aenictinae) From Southeast Asia, pp. 97-102 in Raffles Bulletin of Zoology 61 (1)</i> on page 101, DOI: <a href="http://zenodo.org/record/4509303">10.5281/zenodo.4509303</a&gt

Aenictus levior

Aenictus levior (Karavaivev, 1926): 6 workers from Peninsular Malaysia, Selangor Prov., Ulu Gombak (ca. 250 m alt.), 7 Nov.1999, coll. V. Witte, VW-05 (SKYC); 32 workers from E. Malaysia, Borneo, Sarawak, Mulu, 12 Dec.1993, Sk. Yamane (SKYC, THNHM); 25 workers from E. Malasia, Borneo, Sabah, Logging area near Ranau, 27 Jun.1998, coll. K. Eguchi, Eg98-BOR-841 (SKYC, THNHM); 13 workers from Brunei, Temburong, Kuala Belalong, 19 Feb.1999, K. Eguchi, Eg99-BOR-201 (SKYC, THNHM).Published as part of Jaitrong, Weeyawat & Wiwatwitaya, Decha, 2013, Two New Species Of The Aenictus Pachycerus Species Group (Hymenoptera: Formicidae: Aenictinae) From Southeast Asia, pp. 97-102 in Raffles Bulletin of Zoology 61 (1) on page 101, DOI: 10.5281/zenodo.450930

Aenictus paradentatus Jaitrong & Yamane 2012

<p> <i>Aenictus paradentatus</i> Jaitrong & Yamane, 2012:</p> <p>Holotype and 17 paratype workers from N. Thailand, Chiang Mai Prov., Muang Dist., Doi Suthep-Pui National Park, 20 Aug.1998, coll. W. Jaitrong, WJT98-PD01 (BMNH, MCZC, MHNG, SKYC, THNHM).</p>Published as part of <i>Jaitrong, Weeyawat & Wiwatwitaya, Decha, 2013, Two New Species Of The Aenictus Pachycerus Species Group (Hymenoptera: Formicidae: Aenictinae) From Southeast Asia, pp. 97-102 in Raffles Bulletin of Zoology 61 (1)</i> on page 101, DOI: <a href="http://zenodo.org/record/4509303">10.5281/zenodo.4509303</a&gt

Aenictus nesiotis Wheeler & Chapman 1930

Aenictus nesiotis Wheeler & Chapman in Wheeler, 1930: 21 syntype workers (8 were collected on 4/12/27 and 13 on 11/29/25; collection date was written on the back side of the uppermost label on each pin) from three colonies found at Dumaguete (MCZC).Published as part of Jaitrong, Weeyawat & Wiwatwitaya, Decha, 2013, Two New Species Of The Aenictus Pachycerus Species Group (Hymenoptera: Formicidae: Aenictinae) From Southeast Asia, pp. 97-102 in Raffles Bulletin of Zoology 61 (1) on page 101, DOI: 10.5281/zenodo.450930

Aenictus pachycerus

<i>Aenictus pachycerus</i> species group <p> <b> <i>Diagnosis</i>.</b> — Jaitrong & Yamane (2011) defined this species group as follows: antenna long, consisting of 10 segments; scape long reaching or extending beyond posterolateral corner of head; anterior clypeal margin roundly convex in the middle, lacking denticles; mandible triangular, with very dense punctures; its masticatory margin with a large and sharp apical tooth followed by 4–12 small inconspicuous denticles, which gradually reduce in size toward basal angle of mandible; frontal carinae fused at the level of antennal base to form a single carina, and extending less than half length of head, and well developed anteriorly and poorly developed posteriorly; parafrontal ridge present, reaching less than half length of head; seen in profile its anteriormost part well developed and raised as a subtriangular process; occipital margin forming a collar or carina; promesonotum distinctly convex or very weakly convex dorsally and sloping gradually to propodeum; propodeal junction angulated; declivity of propodeum concave, encircled with a rim; subpetiolar process weakly developed.</p> <p>Head entirely sculptured or smooth and shiny. Petiole and postpetiole densely punctate, at least in Southeast Asian species. First gastral segment entirely smooth and shiny, or rarely superficially shagreened, except the base of the tergite and sternite that has dense small punctures. Body black, dark or reddish brown to light or yellowish brown; typhlatta spot absent.</p> <p> <b> <i>Remarks</i>.</b> — The <i>Aenictus pachycerus</i> group consists of relatively large species in terms of body size (TL 3.20–4.65 mm: 1.80–3.00 mm in smaller species). Wilson (1964) and Jaitrong & Yamane (2011) pointed out that this group is closely related to the <i>A. philippinensis</i> group, but can be distinguished from the latter by the mesonotum not visibly demarcated from the mesopleuron, and the metanotal groove almost absent or indistinct (mesopleuron clearly demarcated from metapleuron by a deep groove and from promesonotum by a distinct carina and metanotal groove relatively deep and distinct in the <i>A. philippinensis</i> group). This species group is also related to the <i>Aenictus hottai</i> group in having developed a frontal carina and parafrontal ridge but can be separated from the latter by the first gastral tergite smooth and shiny and by the weakly developed subpetiolar process (the first gastral tergite densely micropunctate and the subpetiolar process well developed in the latter; see Jaitrong & Yamane, 2011).</p> Key to species of the Southeast Asian <i>Aenictus pachycerus</i> species group based on the worker caste <p>1. Head entirely smooth and shiny; dorsum of mesosoma entirely smooth and shiny.....................................................................2</p> <p> <b>–</b> Head entirely sculptured or partly smooth and shiny; dorsum of mesosoma entirely sculptured or partly smooth and shiny4</p> <p> 2. Promesonotum in profile with clearly convex dorsal outline; propodeum lower than promesonotum; body yellowish brown (Philippines)............................................................. <i>A. powersi</i></p> <p> <b>–</b> Mesosoma dorsally flat or feebly convex; body reddish brown.......................................................................................3</p> <p> 3. Smaller species (HW 0.63–0.65 mm); propodeum in profile with feebly convex dorsal outline; longest pronotal hair 0.25–0.28 mm (Philippines)................................................ <i>A. carolianus</i></p> <p> <b>–</b> Larger species (HW 0.75–0.78 mm); propodeum in profile with straight dorsal outline; longest pronotal hair ca. 0.15 mm (Philippines)................................................................ <i>A. reyesi</i></p> <p> 4. First gastral tergite superficially shagreened (Vietnam, Laos, and Thailand).................................................. <i>A. paradentatus</i></p> <p> <b>–</b> First gastral tergite smooth and shiny.....................................5</p> <p>5. Side of head partly smooth and shiny; dorsal face of pronotum partly shiny..............................................................................6</p> <p> <b>–</b> Side of head entirely sculpturate (punctate or reticulate); dorsal face of pronotum entirely sculptured and opaque..................7</p> <p> 6. Area just outside parafrontal ridge shagreened; vertex reticulate, with sparse standing hairs (less than 12); postpetiole almost as long as petiole (Sulawesi)........ <b> <i>A. sulawesiensis</i>, new species</b> </p> <p> <b>–</b> Area just outside parafrontal ridge with several irregular longitudinal rugulae; vertex finely punctate; vertex with denser standing hairs (more than 15); petiole distinctly longer than petiole (Java)................................................ <b> <i>A.</i> sp. 84 of WJT</b> (see Remarks under Material examined for other species)</p> <p> 7. Propodeal junction in profile with protruding edge that is longer than maximum length of propodeal spiracle, very thin, acute, and far overhanging declivitous face; antennal scape longer (SI 143–152) (Malay Peninsula, Sumatra, Borneo and Java)........................................................................................ <i>A. dentatus</i></p> <p> <b>–</b> Edge of propodeal junction not longer than maximum spiracle width and not overhanging the declivitous face; antennal scape shorter (SI 110 or less than)....................................................8</p> <p> 8. Lateral face of pronotum partly smooth and shiny or superficially shagreened with smooth and shiny interspaces; area just outside parafrontal ridge with 3–5 irregular longitudinal rugulae (Borneo)................................................. <b> <i>A. kutai</i>, new species</b> </p> <p> <b>–</b> Lateral face of pronotum entirely sculpturate and opaque; area just outside parafrontal ridge filely punctate..........................9</p> <p> 9. Apical half of femora superficially reticulate with smooth and shiny bottoms; smaller species (TL 3.5–3.60 mm; HW 0.65–0.68 mm) (Philippines, Sulawesi and Australia)............. <i>A. nesiotis</i></p> <p> <b>–</b> ntire femora finely punctate; larger species (TL 3.65–5.10 mm; HW 0.70–0.98 mm)...............................................................10</p> <p> 10. Petiole sessile; subpetiolar process developed, triangular; ventral outline of postpetiole almost straight or weakly convex; larger species (TL 4.85–5.10 mm; HW 0.90–0.98 mm) (S. China and Vietnam).............................................................. <i>A. bobaiensis</i></p> <p> <b>–</b> Petiole subsessile; subpetiolar process low, its ventral outline convex; ventral outline of postpetiole feebly concave; smaller species (TL 3.65–4.20 mm; HW 0.70–0.80 mm) (Malay Peninsula, Borneo, and Buru Island).......................... <i>A. levior</i></p>Published as part of <i>Jaitrong, Weeyawat & Wiwatwitaya, Decha, 2013, Two New Species Of The Aenictus Pachycerus Species Group (Hymenoptera: Formicidae: Aenictinae) From Southeast Asia, pp. 97-102 in Raffles Bulletin of Zoology 61 (1)</i> on pages 98-99, DOI: <a href="http://zenodo.org/record/4509303">10.5281/zenodo.4509303</a&gt