Spatial connectedness imposes local‐ and metapopulation‐level selection on life history through feedbacks on demography

Dispersal evolution impacts the fluxes of individuals and hence, connectivity in metapopulations. Connectivity is therefore decoupled from the structural connectedness of the patches within the spatial network. Because of demographic feedbacks, local selection also drives the evolution of other life history traits. We investigated how different levels of connectedness affect trait evolution in experimental metapopulations of the two-spotted spider mite. We separated local- and metapopulation-level selection and linked trait divergence to population dynamics. With lower connectedness, an increased starvation resistance and delayed dispersal evolved. Reproductive performance evolved locally by transgenerational plasticity or epigenetic processes. Costs of dispersal, but also changes in local densities and temporal fluctuations herein are found to be putative drivers. In addition to dispersal, demographic traits are able to evolve in response to metapopulation connectedness at both the local and metapopulation level by genetic and/or non-genetic inheritance. These trait changes impact the persistence of spatially structured populations

'Tilting' the Universe with the Landscape Multiverse: The 'Dark' Flow

The theory for the selection of the initial state of the universe from the landscape multiverse predicts superhorizon inhomogeneities induced by nonlocal entanglement of our Hubble volume with modes and domains beyond the horizon. Here we show these naturally give rise to a bulk flow with correlation length of order horizon size. The modification to the gravitational potential has a characteristic scale $L_{1} \simeq 10^{3} H^{-1}$, and it originates from the preinflationary remnants of the landscape. The 'tilt' in the potential induces power to the lowest CMB multipoles, with the dominant contribution being the dipole and next, the quadrupole. The induced multipoles $l \le 2$ are aligned with an axis normal to their alignment plane being oriented along the preferred frame determined by the dipole. The preferred direction is displayed by the velocity field of the bulk flow relative to the expansion frame of the universe. The parameters are tightly constrained thus the derived modifications lead to robust predictions for testing our theory. The 'dark' flow was recently discovered by Kashlinsky et al. to be about $700 km/s$ which seems in good agreement with our predictions for the induced dipole of order $3 \mu K$. Placed in this context, the discovery of the bulk flow by Kashlinsky et al. becomes even more interesting as it may provide a probe of the preinflationary physics and a window onto the landscape multiverse.Comment: 7 pgs, 2 fig

Baryons: What, When and Where?

We review the current state of empirical knowledge of the total budget of baryonic matter in the Universe as observed since the epoch of reionization. Our summary examines on three milestone redshifts since the reionization of H in the IGM, z = 3, 1, and 0, with emphasis on the endpoints. We review the observational techniques used to discover and characterize the phases of baryons. In the spirit of the meeting, the level is aimed at a diverse and non-expert audience and additional attention is given to describe how space missions expected to launch within the next decade will impact this scientific field.Comment: Proceedings Review for "Astrophysics in the Next Decade: JWST and Concurrent Facilities", ed. X. Tielens, 38 pages, 10 color figures. Revised to address comments from the communit

Observations of the High Redshift Universe

(Abridged) In these lectures aimed for non-specialists, I review progress in understanding how galaxies form and evolve. Both the star formation history and assembly of stellar mass can be empirically traced from redshifts z~6 to the present, but how the various distant populations inter-relate and how stellar assembly is regulated by feedback and environmental processes remains unclear. I also discuss how these studies are being extended to locate and characterize the earlier sources beyond z~6. Did early star-forming galaxies contribute significantly to the reionization process and over what period did this occur? Neither theory nor observations are well-developed in this frontier topic but the first results presented here provide important guidance on how we will use more powerful future facilities.Comment: To appear in `First Light in Universe', Saas-Fee Advanced Course 36, Swiss Soc. Astrophys. Astron. in press. 115 pages, 64 figures (see http://www.astro.caltech.edu/~rse/saas-fee.pdf for hi-res figs.) For lecture ppt files see http://obswww.unige.ch/saas-fee/preannouncement/course_pres/overview_f.htm