Measurements of plasma motions in dynamic fibrils

We present a 40 minute time series of filtergrams from the red and the blue wing of the \halpha line in an active region near the solar disk center. From these filtergrams we construct both Dopplergrams and summed ``line center'' images. Several dynamic fibrils (DFs) are identified in the summed images. The data is used to simultaneously measure the proper motion and the Doppler signals in DFs. For calibration of the Doppler signals we use spatially resolved spectrograms of a similar active region. Significant variations in the calibration constant for different solar features are observed, and only regions containing DFs have been used in order to reduce calibration errors. We find a coherent behavior of the Doppler velocity and the proper motion which clearly demonstrates that the evolution of DFs involve plasma motion. The Doppler velocities are found to be a factor 2--3 smaller than velocities derived form proper motions in the image plane. The difference can be explained by the radiative processes involved, the Doppler velocity is a result of the local atmospheric velocity weighted with the response function. As a result the Doppler velocity originates from a wide range in heights in the atmosphere. This is contrasted by the proper motion velocity which is measured from the sharply defined bright tops of the DFs and is therefore a very local velocity measure. The Doppler signal originates from well below the top of the DF. Finally we discuss how this difference together with the lacking spatial resolution of older observations have contributed to some of the confusion about the identity of DFs, spicules and mottles.Comment: 8 pages, 7 figures, Accepted in ApJ, see http://www.astro.uio.no/~oysteol for better quality figures and mpg movi

Spectroscopic measurements of dynamic fibrils in the Ca {\small{II}} 8662 {\AA} line

We present high spatial resolution spectroscopic measurements of dynamic fibrils (DFs) in the Ca {\small{II}} 8662 {\AA} line. These data show clear Doppler shifts in the identified DFs, which demonstrates that at least a subset of DFs are actual mass motions in the chromosphere. A statistical analysis of 26 DFs reveals a strong and statistically significant correlation between the maximal velocity and the deceleration. The range of the velocities and the decelerations are substantially lower, about a factor two, in our spectroscopic observations compared to the earlier results based on proper motion in narrow band images. There are fundamental differences in the different observational methods; when DFs are observed spectroscopically the measured Doppler shifts are a result of the atmospheric velocity, weighted with the response function to velocity over an extended height. When the proper motion of DFs is observed in narrow band images, the movement of the top of the DF is observed. This point is sharply defined because of the high contrast between the DF and the surroundings. The observational differences between the two methods are examined by several numerical experiments using both numerical simulations and a time series of narrow band H$\alpha$ images. With basis in the simulations we conclude that the lower maximal velocity is explained by the low formation height of the Ca IR line. We conclude that the present observations support the earlier result that DFs are driven by magneto-acoustic shocks exited by convective flows and p-modes.Comment: 7 pages 5 figures, Submitted to Ap

Velocities measured in small scale solar magnetic elements

We have obtained high resolution spectrograms of small scale magnetic structures with the Swedish 1-m Solar Telescope. We present Doppler measurements at 0\farcs{2} spatial resolution of bright points, ribbons and flowers and their immediate surroundings, in the C {\small{I}} 5380.3 {\AA} line (formed in the deep photosphere) and the two Fe {\small{I}} lines at 5379.6 {\AA} and 5386.3 {\AA}. The velocity inside the flowers and ribbons are measured to be almost zero, while we observe downflows at the edges. These downflows are increasing with decreasing height. We also analyze realistic magneto-convective simulations to obtain a better understanding of the interpretation of the observed signal. We calculate how the Doppler signal depends on the velocity field in various structures. Both the smearing effect of the non-negligible width of this velocity response function along the line of sight and of the smearing from the telescope and atmospheric point spread function are discussed. These studies lead us to the conclusion that the velocity inside the magnetic elements are really upflow of the order 1--2 km s${}^{-1}$ while the downflows at the edges really are much stronger than observed, of the order 1.5--3.3 km s${}^{-1}$

Stage-specific biomass overcompensation by juveniles in response to increased adult mortality in a wild fish population

Recently developed theoretical models of stage-structured consumer–resource systems have shown that stage-specific biomass overcompensation can arise in response to increased mortality rates. We parameterized a stage-structured population model to simulate the effects of increased adult mortality caused by a pathogen outbreak in the perch (Perca fluviatilis) population of Windermere (UK) in 1976. The model predicts biomass overcompensation by juveniles in response to increased adult mortality due to a shift in food-dependent growth and reproduction rates. Considering cannibalism between life stages in the model reinforces this compensatory response due to the release from predation on juveniles at high mortality rates. These model predictions are matched by our analysis of a 60- year time series of scientific monitoring of Windermere perch, which shows that the pathogen outbreak induced a strong decrease in adult biomass and a corresponding increase in juvenile biomass. Age-specific adult fecundity and size at age were higher after than before the disease outbreak, suggesting that the pathogen-induced mortality released adult perch from competition, thereby increasing somatic and reproductive growth. Higher juvenile survival after the pathogen outbreak due to a release from cannibalism likely contributed to the observed biomass overcompensation. Our findings have general implications for predicting population- and community-level responses to increased size-selective mortality caused by exploitation or disease outbreaks

Six decades of pike and perch population dynamics in Windermere

We use six decades of catch-at-age data for perch (Perca fluviatilis) and pike (Esox lucius) in Windermere (UK) to estimate age- and sex-specific population sizes, natural mortalities and catchabilities in both species. Population sizes are estimated by fitting age structured population models to the catch-at-age data using standard maximum likelihood methods. We validate our methods using data simulations, and use our estimates of vital rates (natural mortality, recruitment and catahability) to address important aspects of fisheries biology. Our model indicates that strong fishery selection against male perch apparently triggered a population collapse, highlighting that sex-selective fisheries can be harmful even at a reasonable exploitation rate (here ≤30%). Recruitment (R) increased with the abundance of spawners (S) in both species, but it also responded to both abiotic and other biotic factors. In particular, increased predator (pike) abundance induced a change from compensation to depensation in the prey (perch) SR relationship, thus favouring the occurrence of an Allee effect. Our study provides reference points for the effective exploitation of pike and perch populations, and underscores the need for ecosystem-based harvesting management

Pathogen-induced rapid evolution in a vertebrate life-history trait

Anthropogenic factors, including climate warming, are increasing the incidence and prevalence of infectious diseases worldwide. Infectious diseases caused by pathogenic parasites can have severe impacts on host survival, thereby altering the selection regime and inducing evolutionary responses in their hosts. Knowledge about such evolutionary consequences in natural populations is critical to mitigate potential ecological and economic effects. However, studies on pathogen-induced trait changes are scarce and the pace of evolutionary change is largely unknown, particularly in vertebrates. Here, we use a time series from long-term monitoring of perch to estimate temporal trends in the maturation schedule before and after a severe pathogen outbreak. We show that the disease induced a phenotypic change from a previously increasing to a decreasing size at maturation, the most important life-history transition in animals. Evolutionary rates imposed by the pathogen were high and comparable to those reported for populations exposed to intense human harvesting. Pathogens thus represent highly potent drivers of adaptive phenotypic evolution in vertebrates