Kinship and its consequences in the cooperatively breeding southern pied babbler Turdoides bicolor

Includes abstract.Includes bibliographical references (p. 119-132).Cooperative breeding occurs when more than two individuals help to raise offspring that are not their own, and has been the focus of empirical and theoretical research for over forty years. Of central importance to this work are the fitness costs and benefits of helping, and the factors limiting the reproduction of helpers. To understand these, the genetic relationship between individuals must be known. In this thesis, I use genetic and observational data to explore kinship between individuals in groups of wild Southern Pied Babblers Turdoides bicolor

Kinship, dear enemies, and costly combat:The effects of relatedness on territorial overlap and aggression in a cooperative breeder

Many species maintain territories, but the degree of overlap between territories and the level of aggression displayed in territorial conflicts can vary widely, even within species. Greater territorial overlap may occur when neighboring territory holders are close relatives. Animals may also differentiate neighbors from strangers, with more familiar neighbors eliciting less‐aggressive responses during territorial conflicts (the “dear enemy” effect). However, research is lacking in how both kinship and overlap affect territorial conflicts, especially in group‐living species. Here, we investigate kinship, territorial overlap, and territorial conflict in a habituated wild population of group‐living cooperatively breeding birds, the southern pied babbler Turdoides bicolor. We find that close kin neighbors are beneficial. Territories overlap more when neighboring groups are close kin, and these larger overlaps with kin confer larger territories (an effect not seen for overlaps with unrelated groups). Overall, territorial conflict is costly, causing significant decreases in body mass, but conflicts with kin are shorter than those conducted with nonkin. Conflicts with more familiar unrelated neighbors are also shorter, indicating these neighbors are “dear enemies.” However, kinship modulates the “dear enemy” effect; even when kin are encountered less frequently, kin elicit less‐aggressive responses, similar to the “dear enemy” effect. Kin selection appears to be a main influence on territorial behavior in this species. Groups derive kin‐selected benefits from decreased conflicts and maintain larger territories when overlapping with kin, though not when overlapping with nonkin. More generally, it is possible that kinship extends the “dear enemy” effect in animal societies

A pair of planets around HD 202206 or a circumbinary planet?

Long-term precise Doppler measurements with the CORALIE spectrograph reveal the presence of a second planet orbiting the solar-type star HD202206. The radial-velocity combined fit yields companion masses of m_2\sini = 17.4 M_Jup and 2.44 M_Jup, semi-major axes of a = 0.83 AU and 2.55 AU, and eccentricities of e = 0.43 and 0.27, respectively. A dynamical analysis of the system further shows a 5/1 mean motion resonance between the two planets. This system is of particular interest since the inner planet is within the brown-dwarf limits while the outer one is much less massive. Therefore, either the inner planet formed simultaneously in the protoplanetary disk as a superplanet, or the outer Jupiter-like planet formed in a circumbinary disk. We believe this singular planetary system will provide important constraints on planetary formation and migration scenarios.Comment: 9 pages, 14 figures, accepted in A&A, 12-May-200

Immune genotypes, immune responses, and survival in a wild bird population

ACKNOWLEDGEMENTS We thank the Tsawout and Tseycum bands for allowing us to conduct research on Mandarte Island, and to the many contributors to long-term monitoring, especially L. Keller, P. Nietlisbach, and J. Krippel. We also thank C. Ritland, A. Miscampbell, and G. Huber for their assistance in the laboratory. All work was conducted under permit of the Canadian Wildlife Service and UBC Animal Care Committee. Funding Information: This study was generously supported by the Natural Sciences and Engineering Research Council of Canada via a Post‐doctoral Fellowship award to MJNF (PDF‐2014–454522) and a Discovery Grant to EAMS.Peer reviewedPublisher PD

The CORALIE survey for southern extra-solar planets IX. A 1.3-day period brown dwarf disguised as a planet

In this article we present the case of HD 41004 AB, a system composed of a K0V star and a 3.7-magnitude fainter M-dwarf companion separated by only 0.5 arcsec. An analysis of CORALIE radial-velocity measurements has revealed a variation with an amplitude of about 50m/s and a periodicity of 1.3days. This radial-velocity signal is consistent with the expected variation induced by the presence a very low mass giant planetary companion to HD 41004 A, whose light dominates the spectra. The radial-velocity measurements were then complemented with a photometric campaign and with the analysis of the bisector of the CORALIE Cross-Correlation Function (CCF). While the former revealed no significant variations within the observational precision of 0.003-0.004 mag (except for an observed flare event), the bisector analysis showed that the line profiles are varying in phase with the radial-velocity. This latter result, complemented with a series of simulations, has shown that we can explain the observations by considering that HD 41004 B has a brown-dwarf companion orbiting with the observed 1.3-day period. If confirmed, this detection represents the first discovery of a brown dwarf in a very short period (1.3-day) orbit around an M dwarf. Finally, this case should be taken as a serious warning about the importance of analyzing the bisector when looking for planets using radial-velocity techniques.Comment: 16 pages, 17 eps figures, A&A in press (Figure 11 not as in original version due to size

Absolute dimensions of eclipsing binaries. XXVIII. BK Pegasi and other F-type binaries: Prospects for calibration of convective core overshoot

We present a detailed study of the F-type detached eclipsing binary BK Peg, based on new photometric and spectroscopic observations. The two components, which have evolved to the upper half of the main-sequence band, are quite different with masses and radii of (1.414 +/- 0.007 Msun, 1.988 +/- 0.008 Rsun) and (1.257 +/- 0.005 Msun, 1.474 +/- 0.017 Rsun), respectively. The 5.49 day period orbit of BK Peg is slightly eccentric (e = 0.053). The measured rotational velocities are 16.6 +/- 0.2 (primary) and 13.4 +/- 0.2 (secondary) km/s. For the secondary component this corresponds to (pseudo)synchronous rotation, whereas the primary component seems to rotate at a slightly lower rate. We derive an iron abundance of [Fe/H] =-0.12 +/- 0.07 and similar abundances for Si, Ca, Sc, Ti, Cr and Ni. Yonsei-Yale and Victoria-Regina evolutionary models for the observed metal abundance reproduce BK Peg at ages of 2.75 and 2.50 Gyr, respectively, but tend to predict a lower age for the more massive primary component than for the secondary. We find the same age trend for three other upper main-sequence systems in a sample of well studied eclipsing binaries with components in the 1.15-1.70 Msun range, where convective core overshoot is gradually ramped up in the models. We also find that the Yonsei-Yale models systematically predict higher ages than the Victoria-Regina models. The sample includes BW Aqr, and as a supplement we have determined a [Fe/H] abundance of -0.07 +/- 0.11 for this late F-type binary. We propose to use BK Peg, BW Aqr, and other well-studied 1.15-1.70 Msun eclipsing binaries to fine-tune convective core overshoot, diffusion, and possibly other ingredients of modern theoretical evolutionary models.Comment: Accepted for publication in Astronomy and Astrophysic

Paleocurrent reconstruction of the deep Pacific inflow during the middle Miocene : reflections of East Antarctic Ice Sheet growth

Today the deep western boundary current (DWBC) east of New Zealand is the most important route for deep water entering the Pacific Ocean. Large-scale changes in deep water circulation patterns are thought to have been associated with the development of the East Antarctic Ice Sheet (EAIS) close to the main source of bottom water for the DWBC. Here we reconstruct the changing speed of the southwest Pacific DWBC during the middle Miocene from ∼15.5-12.5 Ma, a period of significant global ice accumulation associated with EAIS growth. Sortable silt mean grain sizes from Ocean Drilling Program Site 1123 reveal variability in the speed of the Pacific inflow on the timescale of the 41 kyr orbital obliquity cycle. Similar orbital period flow changes have recently been demonstrated for the Pleistocene epoch. Collectively, these observations suggest that a strong coupling between changes in the speed of the deep Pacific inflow and high-latitude climate forcing may have been a persistent feature of the global thermohaline circulation system for at least the past 15 Myr. Furthermore, long-term changes in flow speed suggest an intensification of the DWBC under an inferred increase in Southern Component Water production. This occurred at the same time as decreasing Tethyan outflow and major EAIS growth between ∼15.5 and 13.5 Ma. These results provide evidence that a major component of the deep thermohaline circulation was associated with the middle Miocene growth of the EAIS and support the view that this time interval represents an important step in the development of the Neogene icehouse climate

Four-colour photometry of eclipsing binaries. XL, uvby light curves for the B-type systems DW Carinae, BF Centauri, AC Velorum, and NSV 5783

Aims. In order to increase the limited number of B-stars with accurately known dimensions, and also the number of well studied eclipsing binaries in open clusters, we have undertaken observations and studies of four southern double-lined eclipsing B-type binaries; DWCar, BF Cen, ACVel, and NSV 5783. Methods. Complete uvby light curves were observed between January 1982 and April 1991 at the Danish 0.5 m telescope at ESO La Silla, since 1985 known as the Strömgren Automatic Telescope (SAT). Standard indices for the systems and the comparison stars,as well as additional minima observations for ACVel, have been obtained later at SAT. For DWCar and ACVel, high-resolution spectra for definitive spectroscopic orbits have also been obtained; they are presented as part of the detailed analyses of these systems. A few spectra of NSV 5783 are included in the present paper. Results. For all four systems, the first modern accurate light curves have been established. DWCar is a detached system consisting of two nearly identical components. It is member of the young open cluster Cr228. A detailed analysis, based on the new light curves and 29 high-resolution spectra, is published separately. BFCen is semidetached and is member of NGC 3766. Modern spectra are needed for a detailed study. ACVel is a detached system with at least one more star. A full analysis, based on the new light curves and 18 high-resolution spectra, is published separately. NSV 5783 is discovered to be an eclipsing binary consisting of two well-detached components in an 11-day period eccentric (e = 0.18) orbit. Secondary eclipse is practically total. From the light curves and a few high-resolution spectra, accurate photometric elements and preliminary absolute dimensions have been determined. The quite similar components have masses of about 5 M and radii of about 3.5 R, and they seem to have evolved just slightly off the ZAMS. The measured rotational velocities (≈150 km s−1) are about 6 times those corresponding to pseudosynchronization

Absolute dimensions of eclipsing binaries. XXVI, Setting a new standard : masses, radii, and abundances for the F-type systems AD Bootis, VZ Hydrae, and WZ Ophiuchi

Context. Accurate mass, radius, and abundance determinations from binaries provide important information on stellar evolution, fundamental to central fields in modern astrophysics and cosmology. Aims. We aim to determine absolute dimensions and abundances for the three F-type main-sequence detached eclipsing binaries ADBoo, VZHya, and WZOph and to perform a detailed comparison with results from recent stellar evolutionary models. Methods. uvby light curves and uvbyβ standard photometry were obtained with the Strömgren Automatic Telescope at ESO, La Silla, radial velocity observations at CfA facilities, and supplementary high-resolution spectra with ESO’s FEROS spectrograph. State-ofthe-art methods were applied for the analyses: the EBOP andWilson-Devinney binary models, two-dimensional cross-correlation and disentangling, and the VWA abundance analysis tool. Results. Masses and radii that are precise to 0.5–0.7% and 0.4–0.9%, respectively, have been established for the components, which span the ranges of 1.1 to 1.4 M and 1.1 to 1.6 R. The [Fe/H] abundances are from –0.27 to +0.10, with uncertainties between 0.07 and 0.15 dex. We find indications of a slight α-element overabundance of [α/Fe] ∼ +0.1 for WZOph. The secondary component of ADBoo and both components of WZOph appear to be slightly active. Yale-Yonsai and Victoria-Regina evolutionary models fit the components of ADBoo and VZHya almost equally well, assuming coeval formation, at ages of about 1.75/1.50 Gyr (ADBoo) and 1.25/1.00 Gyr (VZHya). BaSTI models, however, predict somewhat different ages for the primary and secondary components. For WZOph, the models from all three grids are significantly hotter than observed. A low He content, decreased envelope convection coupled with surface activity, and/or higher interstellar absorption would remove the discrepancy, but its cause has not been definitively identified. Conclusions. We have demonstrated the power of testing and comparing recent stellar evolutionary models using eclipsing binaries, provided their abundances are known. The strongest limitations and challenges are set by Teff and interstellar absorption determinations, and by their effects on and correlation with abundance results

Disk, merger, or outflow ? Molecular gas kinematics in two powerful obscured QSOs at z>3.4

We report on the detection of bright CO(4-3) line emission in two powerful, obscured quasars discovered in the SWIRE survey, SW022513 and SW022550 at z>3.4. We analyze the line strength and profile to determine the gas mass, dynamical mass and the gas dynamics for both galaxies. In SW022513 we may have found the first evidence for a molecular, AGN-driven wind in the early Universe. The line profile in SW022513 is broad (FWHM = 1000 km/s) and blueshifted by -200 km/s relative to systemic (where the systemic velocity is estimated from the narrow components of ionized gas lines, as is commonly done for AGN at low and high redshifts). SW022550 has a more regular, double-peaked profile, which is marginally spatially resolved in our data, consistent with either a merger or an extended disk. The molecular gas masses, 4x10^10 Msun, are large and account for <30% of the stellar mass, making these obscured QSOs as gas rich as other powerful CO emitting galaxies at high redshift, i.e., submillimeter galaxies. Our sources exhibit relatively lower star-formation efficiencies compared to other dusty, powerful starburst galaxies at high redshift. We speculate that this could be a consequence of the AGN perturbing the molecular gas.Comment: Accepted for publication in A&