VADER: A Flexible, Robust, Open-Source Code for Simulating Viscous Thin Accretion Disks

The evolution of thin axisymmetric viscous accretion disks is a classic problem in astrophysics. While models based on this simplified geometry provide only approximations to the true processes of instability-driven mass and angular momentum transport, their simplicity makes them invaluable tools for both semi-analytic modeling and simulations of long-term evolution where two- or three-dimensional calculations are too computationally costly. Despite the utility of these models, the only publicly-available frameworks for simulating them are rather specialized and non-general. Here we describe a highly flexible, general numerical method for simulating viscous thin disks with arbitrary rotation curves, viscosities, boundary conditions, grid spacings, equations of state, and rates of gain or loss of mass (e.g., through winds) and energy (e.g., through radiation). Our method is based on a conservative, finite-volume, second-order accurate discretization of the equations, which we solve using an unconditionally-stable implicit scheme. We implement Anderson acceleration to speed convergence of the scheme, and show that this leads to factor of $\sim 5$ speed gains over non-accelerated methods in realistic problems, though the amount of speedup is highly problem-dependent. We have implemented our method in the new code Viscous Accretion Disk Evolution Resource (VADER), which is freely available for download from https://bitbucket.org/krumholz/vader/ under the terms of the GNU General Public License.Comment: 58 pages, 13 figures, accepted to Astronomy & Computing; this version includes more discussion, but no other changes; code is available for download from https://bitbucket.org/krumholz/vader

Age-dependent induction of immunity and subsequent survival costs in males and females of a temperate damselfly

BACKGROUND: To understand variation in resistance to parasites within host populations, researchers have examined conditions under which immunity is induced and/or is costly. Both host sex and age have been found to influence immune expression and subsequently are likely factors influencing the costs of resistance. The purpose of this study was to examine immune expression and associated survival costs for two age groups (newly emerged and sexually mature individuals) of the damselfly, Enallagma boreale Selys. Survival was assessed for experimentally challenged and control damselflies, housed initially at 22°C and then subjected to low temperatures (15°C) associated with reduced foraging activity and food deprivation. Experimental conditions emulated natural local variation in bouts of good weather followed by inclement weather (successions of days with hourly mean temperatures around 15°C and/or rainy weather). RESULTS: At least one of three immune traits was induced to higher levels for both newly emerged and mature E. boreale challenged by Lippopolysaccharide (LPS) relative to saline-injected controls, when housed at 22°C. The immune traits assayed included haemocyte concentration, Phenoloxidase activity and antibacterial activity and their induction varied among ages and between males and females. For matures, those injected with LPS had lowered survivorship compared to saline-injected controls that were housed initially at 22°C and subsequently at 15°C. Newly emerged LPS-injected damselflies did not show reduced survivorship relative to newly-emerged controls, despite showing immune induction. CONCLUSION: Reduced longevity following induction of immunity was observed for reproductively mature damselflies, but not for newly emerged damselflies. Costs of resistance depend only partly on the immune trait induced and more on the age (but not sex) of the host. In four years, we often observed bouts of inclement weather following good days and these bouts occurred primarily during the emergence periods, but also during the flight periods, of E. boreale. The duration of these bouts appear sufficient to compromise survival of mature damselflies that responded immunologically to LPS challenge. We further suggest the environmental conditions likely experienced by different ages of damselflies, following resistance expression, has influenced optimal immune investment by individuals in different age classes and the likelihood of detecting costs of resistance

Inter-annual variation in prevalence and intensity of mite parasitism relates to appearance and expression of damselfly resistance

<p>Abstract</p> <p>Background</p> <p>Insects can resist parasites using the costly process of melanotic encapsulation. This form of physiological resistance has been studied under laboratory conditions, but the abiotic and biotic factors affecting resistance in natural insect populations are not well understood. Mite parasitism of damselflies was studied in a temperate damselfly population over seven seasons to determine if melanotic encapsulation of mite feeding tubes was related to degree of parasitism, host sex, host size, emergence timing, duration of the emergence period, and average daily air temperature.</p> <p>Results</p> <p>Although parasite prevalence in newly emerged damselflies was > 77% each year, hosts did not resist mites in the early years of study. Resistance began the year that there was a dramatic increase in the number of mites on newly emerged damselflies. Resistance continued to be correlated with mite prevalence and intensity throughout the seven-year study. However, the percentage of hosts resisting only ranged from 0-13% among years and resistance was not sex-biased and was not correlated with host size. Resistance also was not correlated with air temperature or with timing or duration of damselfly emergence.</p> <p>Conclusions</p> <p>Resistance in host damselflies was weakly and variably expressed over the study period. Factors such as temperature, which have been identified in laboratory studies as contributing to resistance by similar hosts, can be irrelevant in natural populations. This lack of temperature effect may be due to the narrow range in temperatures observed at host emergence among years. Degree of mite parasitism predicted both the appearance and continued expression of resistance among parasitized damselflies.</p

Factors Influencing Colony Attendance by Northern Fulmars in the Canadian Arctic

The northern fulmar (Fulmarus glacialis) is a common seabird of the North Atlantic Ocean and marine areas of eastern Nunavut. We studied attendance patterns of fulmars at two colonies in the Canadian High Arctic to assess variation in attendance at high latitudes, to compare these patterns to those found for fulmars breeding at colonies to the south, and to determine how weather conditions and breeding status might influence attendance. Colony attendance patterns shared similarities across the species’ range, although local variation in the timing of food supplies may explain differences in the seasonal timing of peak counts at colonies. Stronger winds and extreme inclement weather resulted in lower counts of birds at colonies. Seasonal and daily variation in attendance was attributable to differences in the numbers of failed breeders or non-breeding birds. Changes toward earlier breakup and reduced extent of sea ice should lead to earlier peaks and continued short periodicities in colony attendance cycles of Arctic fulmars.Le fulmar boréal (Fulmarus glacialis) est un oiseau marin courant du nord de l’océan Atlantique et des régions marines de l’est du Nunavut. Nous avons étudié les tendances de fréquentation des fulmars à deux colonies situées dans l’Extrême-Arctique canadien dans le but d’évaluer les variations de fréquentation en hautes latitudes, de comparer ces tendances à celles caractérisant les fulmars se reproduisant aux colonies se trouvant plus au sud et de déterminer en quoi les conditions météorologiques et le statut de reproduction peuvent exercer une influence sur la fréquentation. Il existait des similarités entre la répartition des espèces sur le plan des tendances de fréquentation au sein des colonies, bien que les variations locales en matière de disponibilité des approvisionnements alimentaires puissent expliquer les différences entre les dénombrements saisonniers de pointe aux colonies. Les vents violents et le temps des plus incléments se sont traduits par des dénombrements d’oiseaux moins grands aux colonies. Les variations de fréquentation saisonnières et quotidiennes étaient attribuables aux différences sur le plan du nombre de reproducteurs manqués ou d’oiseaux ne se reproduisant pas. Les changements qui se manifestent, tels que l’avancement de la débâcle et une moins grande surface de glace de mer, devraient entraîner des pointes plus hâtives et des périodicités continuellement courtes pour ce qui est des cycles de fréquentation des colonies de fulmars boréaux