Can we characterise ‘openness’ in the Holocene palaeoenvironmental record? Modern analogue studies of insect faunas and pollen spectra from Dunham Massey deer park and Epping Forest, England

This paper examines the degree to which tree-associated Coleoptera (beetles) and pollen could be used to predict the degree of ‘openness’ in woodland. The results from two modern insect and pollen analogue studies from ponds at Dunham Massey, Cheshire and Epping Forest, Greater London are presented. We explore the reliability of modern pollen rain and sub-fossil beetle assemblages to represent varying degrees of canopy cover for up to 1000 m from a sampling site. Modern woodland canopy structure around the study sites has been assessed using GIS-based mapping at increasing radial distances as an independent check on the modern insect and pollen data sets. These preliminary results suggest that it is possible to use tree-associated Coleoptera to assess the degree of local vegetation openness. In addition, it appears that insect remains may indicate the relative intensity of land use by grazing animals. Our results also suggest most insects are collected from within a 100—200 m radius of the sampling site. The pollen results suggest that local vegetation and density of woodland in the immediate area of the sampling site can have a strong role in determining the pollen signal

Pinus and Prostomis: a dendrochronological and palaeoentomological study of a study of a mid-Holocene woodland in Eastern England

Tree-ring analysis of subfossil Pinus sylvestris L. and Quercus sp. and their associated subfossil insect assemblages from tree rot-holes have been used to study a prehistoric forest buried in the basal peats at Tyrham Hall Quarry, Hatfield Moors SSSI, in the Humberhead Levels, eastern England. The site provided a rare opportunity to examine the date, composition, age structure and entomological biodiversity of a mid-Holocene Pinus-dominated forest. The combined approaches of dendrochronology and palaeoentomology have enabled a detailed picture of the forest to be reconstructed, within a precise time-frame. The Pinus chronology has been precisely dated to 2921–2445 bc against the English Quercus master curve and represents the first English Pinus chronology to be dendrochronologically dated. A suite of important xylophilous (wood-loving) beetles that are today very rare and four species that no longer live within the British Isles were also recovered, their disappearance associated with the decline in woodland habitats as well as possible climatic change. The subfossil insects indicate that the characteristic species of the site's modern-day fauna were already in place 4000 years ago. These findings have important implications in terms of maintaining long-term invertebrate biodiversity of forest and mire sites

Archaeological and environmental investigations of Late Glacial and Holocene river valley sequences on the River Soar, at Croft, Leicestershire

The sediments, stratigraphy and archaeology of several sections through Lateglacial and Holocene deposits associated with the past course of the Thurlston Brook at Croft, Leicestershire, UK are outlined. The results of pollen, plant macrofossil and insect analyses from these deposits are presented and this information is used to provide a detailed reconstruction of changing river conditions and human land use at this location during the Holocene. Despite the presence of hiatuses in the record seen at Croft, and other sites of this scale, with accurate work and clear dating controls it is possible to 'stitch' together continuous detailed sequences. The general pattern of Holocene landscape and fluvial change appears to echo that seen in the Trent valley region and nationally. It is suggested that small headwater catchments such as at Croft have the potential to provide detailed and sensitive records of Holocene events comparable with those from 'standard' sediment sequences

Mire ontogeny, environmental and climate change inferred from fossil beetle successions from Hatfield Moors, eastern England

Results of a fossil Coleoptera (beetle) fauna from a fen edge sequence from Hatfield Moors, Humber-head Levels, England, are presented. Mire ontogeny inferred from this location and others are discussed, particularly in the light of previous palynological and plant macrofossil investigations. Peat initiation across most of the site centres around 3000 cal. bc, characterized by a Calluna-Eriophorum heath with areas of PinusBetula woodland. The onset of peat accumulation on the southern margins of the site was delayed until 1520– 1390 cal. bc and appears to overlap closely with a recurrence surface at a pollen site (HAT 2) dated to 1610– 1440 cal. bc, suggesting that increased surface wetness may have caused mire expansion at this time. The faunas illustrate the transition from eutrophic and mesotrophic fen to ombrotrophic raised mire, although the significance of both Pinus- and Calluna-indicating species through the sequence suggests that heath habitats may have continued to be important. Elsewhere, this earlier phase of rich fen is lacking and mesotrophic mire developed immediately above nutrient-poor sands, with ombrotrophic conditions indicated soon after. Correspondence analysis of the faunas provides valuable insights into the importance of sandy heath habitats on Hatfield Moors. The continuing influence of the underlying coversands suggests these may have been instrumental in mire ontogeny. The research highlights the usefulness of using Coleoptera to assess mire ontogeny, fluctuations in site hydrology and vegetation cover, particularly when used in conjunction with other peatland proxies. The significance of a suite of extinct beetle species is discussed with reference to forest history and climatic change