Apparent slip due to the motion of suspended particles in flows of electrolyte solutions

We consider pressure-driven flows of electrolyte solutions in small channels or capillaries in which tracer particles are used to probe velocity profiles. Under the assumption that the double layer is thin compared to the channel dimensions, we show that the flow-induced streaming electric field can create an apparent slip velocity for the motion of the particles, even if the flow velocity still satisfies the no-slip boundary condition. In this case, tracking of particle would lead to the wrong conclusion that the no-slip boundary condition is violated. We evaluate the apparent slip length, compare with experiments, and discuss the implications of these results

A note on the stability of slip channel flows

We consider the influence of slip boundary conditions on the modal and non-modal stability of pressure-driven channel flows. In accordance with previous results by Gersting (1974) (Phys. Fluids, 17) but in contradiction with the recent investigation of Chu (2004) (C.R. Mecanique, 332), we show that slip increases significantly the value of the critical Reynolds number for linear instability. The non-modal stability analysis however reveals that the slip has a very weak influence on the maximum transient energy growth of perturbations at subcritical Reynolds numbers. Slip boundary conditions are therefore not likely to have a significant effect on the transition to turbulence in channel flows

Phoretic self-propulsion at finite P\'eclet numbers

Phoretic self-propulsion is a unique example of force- and torque-free motion on small scales. The classical framework describing the flow field around a particle swimming by self-diffusiophoresis neglects the advection of the solute field by the flow and assumes that the chemical interaction layer is thin compared to the particle size. In this paper we quantify and characterize the effect of solute advection on the phoretic swimming of a sphere. We first rigorously derive the regime of validity of the thin-interaction layer assumption at finite values of the P\'eclet number (Pe). Within this assumption, we solve computationally the flow around Janus phoretic particles and examine the impact of solute advection on propulsion and the flow created by the particle. We demonstrate that although advection always leads to a decrease of the swimming speed and flow stresslet at high values of the P\'eclet number, an increase can be obtained at intermediate values of Pe. This possible enhancement of swimming depends critically on the nature of the chemical interactions between the solute and the surface. We then derive an asymptotic analysis of the problem at small Pe allowing to rationalize our computational results. Our computational and theoretical analysis is accompanied by a parallel study of the role of reactive effects at the surface of the particle on swimming (Damk\"ohler number).Comment: 27 pages, 15 figures, to appear in J. Fluid Mec

Life around the scallop theorem

Locomotion on small scales is dominated by the effects of viscous forces and, as a result, is subject to strong physical and mathematical constraints. Following Purcell's statement of the scallop theorem which delimitates the types of swimmer designs which are not effective on small scales, we review the different ways the constraints of the theorem can be escaped for locomotion purposes

Autophoretic locomotion from geometric asymmetry

Among the few methods which have been proposed to create small-scale swimmers, those relying on self-phoretic mechanisms present an interesting design challenge in that chemical gradients are required to generate net propulsion. Building on recent work, we propose that asymmetries in geometry are sufficient to induce chemical gradients and swimming. We illustrate this idea using two different calculations. We first calculate exactly the self-propulsion speed of a system composed of two spheres of unequal sizes but identically chemically homogeneous. We then consider arbitrary, small-amplitude, shape deformations of a chemically-homogeneous sphere, and calculate asymptotically the self-propulsion velocity induced by the shape asymmetries. Our results demonstrate how geometric asymmetries can be tuned to induce large locomotion speeds without the need of chemical patterning.Comment: 17 pages, 10 figure

Efficiency optimization and symmetry-breaking in a model of ciliary locomotion

A variety of swimming microorganisms, called ciliates, exploit the bending of a large number of small and densely-packed organelles, termed cilia, in order to propel themselves in a viscous fluid. We consider a spherical envelope model for such ciliary locomotion where the dynamics of the individual cilia are replaced by that of a continuous overlaying surface allowed to deform tangentially to itself. Employing a variational approach, we determine numerically the time-periodic deformation of such surface which leads to low-Reynolds locomotion with minimum rate of energy dissipation (maximum efficiency). Employing both Lagrangian and Eulerian points of views, we show that in the optimal swimming stroke, individual cilia display weak asymmetric beating, but that a significant symmetry-breaking occurs at the organism level, with the whole surface deforming in a wave-like fashion reminiscent of metachronal waves of biological cilia. This wave motion is analyzed using a formal modal decomposition, is found to occur in the same direction as the swimming direction, and is interpreted as due to a spatial distribution of phase-differences in the kinematics of individual cilia. Using additional constrained optimizations, as well as a constructed analytical ansatz, we derive a complete optimization diagram where all swimming efficiencies, swimming speeds, and amplitude of surface deformation can be reached, with the mathematically optimal swimmer, of efficiency one half, being a singular limit. Biologically, our work suggests therefore that metachronal waves may allow cilia to propel cells forward while reducing the energy dissipated in the surrounding fluid.Comment: 29 pages, 20 figure

Unsteady feeding and optimal strokes of model ciliates

The flow field created by swimming microorganisms not only enables their locomotion but also leads to advective transport of nutrients. In this paper we address analytically and computationally the link between unsteady feeding and unsteady swimming on a model microorganism, the spherical squirmer, actuating the fluid in a time-periodic manner. We start by performing asymptotic calculations at low P\'eclet number (Pe) on the advection-diffusion problem for the nutrients. We show that the mean rate of feeding as well as its fluctuations in time depend only on the swimming modes of the squirmer up to order Pe^(3/2), even when no swimming occurs on average, while the influence of non-swimming modes comes in only at order Pe^2. We also show that generically we expect a phase delay between feeding and swimming of 1/8th of a period. Numerical computations for illustrative strokes at finite Pe confirm quantitatively our analytical results linking swimming and feeding. We finally derive, and use, an adjoint-based optimization algorithm to determine the optimal unsteady strokes maximizing feeding rate for a fixed energy budget. The overall optimal feeder is always the optimal steady swimmer. Within the set of time-periodic strokes, the optimal feeding strokes are found to be equivalent to those optimizing periodic swimming for all values of the P\'eclet number, and correspond to a regularization of the overall steady optimal.Comment: 26 pages, 11 figures, to appear in Journal of Fluid Mechanic