On the impact of dispersal asymmetry on metapopulation persistence

Metapopulation theory for a long time has assumed dispersal to be symmetric, i.e. patches are connected through migrants dispersing bi-directionally without a preferred direction. However, for natural populations symmetry is often broken, e.g. for species in the marine environment dispersing through the transport of pelagic larvae with ocean currents. The few recent studies of asymmetric dispersal concluded, that asymmetry has a distinct negative impact on the persistence of metapopulations. Detailed analysis however revealed, that these previous studies might have been unable to properly disentangle the effect of symmetry from other potentially confounding properties of dispersal patterns. We resolve this issue by systematically investigating the symmetry of dispersal patterns and its impact on metapopulation persistence. Our main analysis based on a metapopulation model equivalent to previous studies but now applied on regular dispersal patterns aims to isolate the effect of dispersal symmetry on metapopulation persistence. Our results suggest, that asymmetry in itself does not imply negative effects on metapopulation persistence. For this reason we recommend to investigate it in connection with other properties of dispersal instead of in isolation.Comment: 19 pages, 5 figure

Estimation of drift and diffusion functions from time series data: A maximum likelihood framework

Complex systems are characterized by a huge number of degrees of freedom often interacting in a non-linear manner. In many cases macroscopic states, however, can be characterized by a small number of order parameters that obey stochastic dynamics in time. Recently techniques for the estimation of the corresponding stochastic differential equations from measured data have been introduced. This contribution develops a framework for the estimation of the functions and their respective (Bayesian posterior) confidence regions based on likelihood estimators. In succession approximations are introduced that significantly improve the efficiency of the estimation procedure. While being consistent with standard approaches to the problem this contribution solves important problems concerning the applicability and the accuracy of estimated parameters.Comment: 18 pages, 2 figure

An assessment of community-based social enterprise in three European countries

See pages 2-7 of the report

The implications of Schumpeter's theories of innovation for the role, organisation and impact of community-based social enterprise in three European countries

Social enterprises, with strong ties to local areas and communities, have been a growing phenomenon in many European countries at least since the financial crisis of 2007-2008 and the resulting retrenchment of state involvement in welfare provision. The paper draws on the empirical findings from nine case studies of community-based social enterprises (CBSE) in three countries which were investigated in depth in our study. Our objective is to use Schumpeter’s work as a lens to assess the effects of social innovation on different aspects of this type of social organisation. Thus, we aim to address the questions: (i) to what extent can CBSEs be considered as a form of social innovation and how does this innovation arise in terms of role, organisation and impact of CB- SEs? (ii) What are the similarities and differences between CBSEs in the three selected European countries? And (iii) how far does Schumpeter’s conceptual framework of “creative destruction” provide insights into the process of organisational change in this form of social enterprise? In do- ing so we identify and discuss a series of innovations in organisation, project selection and delivery and conclude with insights relating to Schumpeter’s theory of “creative destruction”

Groundwater seepage landscapes from distant and local sources in experiments and on Mars

© 2014 Author(s). Valleys with theater-shaped heads can form due to the seepage of groundwater and as a result of knickpoint (waterfall) erosion generated by overland flow. This ambiguity in the mechanism of formation hampers the interpretation of such valleys on Mars, particularly since there is limited knowledge of material properties. Moreover, the hydrological implications of a groundwater or surface water origin are important for our understanding of the evolution of surface features on Mars, and a quantification of valley morphologies at the landscape scale may provide diagnostic insights on the formative hydrological conditions. However, flow patterns and the resulting landscapes produced by different sources of groundwater are poorly understood. We aim to improve the understanding of the formation of entire valley landscapes through seepage processes from different groundwater sources that will provide a framework of landscape metrics for the interpretation of such systems. We study groundwater seepage from a distant source of groundwater and from infiltration of local precipitation in a series of sandbox experiments and combine our results with previous experiments and observations of the Martian surface. Key results are that groundwater flow piracy acts on valleys fed by a distant groundwater source and results in a sparsely dissected landscape of many small and a few large valleys. In contrast, valleys fed by a local groundwater source, i.e., nearby infiltration, result in a densely dissected landscape. In addition, valleys fed by a distant groundwater source grow towards that source, while valleys with a local source grow in a broad range of directions and have a strong tendency to bifurcate, particularly on flatter surfaces. We consider these results with respect to two Martian cases: Louros Valles shows properties of seepage by a local source of groundwater and Nirgal Vallis shows evidence of a distant source, which we interpret as groundwater flow from Tharsis