Plagiochila rutilans (Hepaticae): A poorly known species from tropical America

The neotropical liverwort, Plagiochila rutilans Lindenb., is conspecific with P. remotifolia Hampe and Gottsche, P. farlowii Steph., P. harrisana Steph, and P. organensis Herzog. Plagiochila standleyi Carl is reduced to a variety of P. rutilans. Plagiochila gymnocalycina (Lehm. and Lindenb.) Mont. and P. portoricensis Hampe and Gottsche (= P. simplex (Sw.) Lindenb.) are excluded from the synonymy of P. rutilans. Plagiochila rutilans var. liebmanniana Gottsche is a synonym of P. crispabilis Lindenb.; P. rutilans var. laxa Lindenb. and var. angustifolia Herzog are conspecific with P. gymnocalycina. Sporophytes of P. rutilans are described for the first time. Fresh material of P. rutilans exhibits a distinct odor of peppermint caused by the presence of several menthane monoterpenoids, principally pulegone. NMR (nuclear magnetic resonance) fingerprints and GC-MS data indicate that the lipophilic secondary metabolite profiles are distinct for the two varieties accepted in this study