The multi-peak adaptive landscape of crocodylomorph body size evolution

Background: Little is known about the long-term patterns of body size evolution in Crocodylomorpha, the > 200-million-year-old group that includes living crocodylians and their extinct relatives. Extant crocodylians are mostly large-bodied (3–7 m) predators. However, extinct crocodylomorphs exhibit a wider range of phenotypes, and many of the earliest taxa were much smaller ( Results: Crocodylomorphs reached an early peak in body size disparity during the Late Jurassic, and underwent an essentially continual decline since then. A multi-peak Ornstein-Uhlenbeck model outperforms all other evolutionary models fitted to our data (including both uniform and non-uniform), indicating that the macroevolutionary dynamics of crocodylomorph body size are better described within the concept of an adaptive landscape, with most body size variation emerging after shifts to new macroevolutionary regimes (analogous to adaptive zones). We did not find support for a consistent evolutionary trend towards larger sizes among lineages (i.e., Cope’s rule), or strong correlations of body size with climate. Instead, the intermediate to large body sizes of some crocodylomorphs are better explained by group-specific adaptations. In particular, the evolution of a more aquatic lifestyle (especially marine) correlates with increases in average body size, though not without exceptions. Conclusions: Shifts between macroevolutionary regimes provide a better explanation of crocodylomorph body size evolution on large phylogenetic and temporal scales, suggesting a central role for lineage-specific adaptations rather than climatic forcing. Shifts leading to larger body sizes occurred in most aquatic and semi-aquatic groups. This, combined with extinctions of groups occupying smaller body size regimes (particularly during the Late Cretaceous and Cenozoic), gave rise to the upward-shifted body size distribution of extant crocodylomorphs compared to their smaller-bodied terrestrial ancestors.</p

Human rights, Islam and the failure of cosmopolitanism

The rise of global human rights has been presented as compelling evidence for cosmopolitan progress, especially in Europe, and with particular benefits for ethnic and religious minorities. New conceptions of citizenship – post-national, de-nationalized, disaggregated and cosmopolitan – have been used to show how minorities have created and profited from European cosmopolitanism. Some theorists have pointed to human rights activism, especially around the foulard affair, to illustrate the arrival of cosmopolitan justice. However, this paper suggests that cosmopolitan optimism has misjudged the magnitude of the impact of human rights. European cosmopolitanism’s commitment to ‘cool’ attachments has difficulty with ‘thick’ religious attachments. Muslim cosmopolitanism – expressed for example though religious pilgrimages – makes Muslims ‘bad’ cosmopolitans in the European version. This clash needs to be reconciled before Europe can define itself as the unrivalled source of cosmopolitan justice