Effect of stand-replacing fires on Mediterranean plant species in their marginal alpine range

In the southern side of the Alps, many relic species with Mediterranean and sub-Mediterranean distribution were described in mild-winter, fire-prone areas. Very few studies have modeled the importance of environmental factors on their distribution. In this paper, we assessed the effect of fire on the occurrence of euri- and steno-Mediterranean (ESM) species in Pinus sylvestris forests of Aosta Valley (Italy), by analyzing vegetation in a chronosequence of six stand-replacing fires (1962-2006). We analyzed species richness along the chronosequence, and modeled it as a function of time since fire, environment, and stand structure. We observed a strong positive association between ESM and total species richness. Temporal vegetation dynamics did not follow the direct succession pattern that is commonly observed in Mediterranean ecosystems. Two distinct maxima of ESM species richness were observed: (1) short lived, ruderal species (32 % of all ESM species) in the early post-fire stages, and (2) dry grassland species (54 %) in intermediate stages. The first were facilitated by the consumption of canopy and litter during fire, while the second by delayed tree canopy closure. In multivariate models of ESM species richness, light and elevation were the only significant predictors. Contrary to expectations, time since fire was not significant. Our study suggests that stand-replacing fires play an important role in preserving Mediterranean species in the study area by maintaining an open canopy, and promote local species diversity

Scaling of the Critical Function for the Standard Map: Some Numerical Results

The behavior of the critical function for the breakdown of the homotopically non-trivial invariant (KAM) curves for the standard map, as the rotation number tends to a rational number, is investigated using a version of Greene's residue criterion. The results are compared to the analogous ones for the radius of convergence of the Lindstedt series, in which case rigorous theorems have been proved. The conjectured interpolation of the critical function in terms of the Bryuno function is discussed.Comment: 26 pages, 3 figures, 13 table

Stand and coarse woody debris dynamics in subalpine Norway spruce forests withdrawn from regular management

We studied structural characteristics, amount and quality of coarse woody debris (CWD), intensity of competition and mortality in two subalpine Norway spruce stands withdrawn from regular management. The stands, that we measured twice (in 1993 and 2005), have similar age and structure, but a different time has elapsed since the last silvicultural treatments (respectively 22 and about 55 y). The main purposes were to analyze the current stage of development as compared to the old-growth one and to highlight the legacies of past management. Although relatively old, the first plot (Valbona 1) was at the end of the pole stage. CWD was low in volume and was mainly of man-made origin (stumps). A recent thinning from below has reduced density-dependent competition and delayed the development of old-growth characteristics. The second plot (Valbona 2a) was at the beginning of the transition stage, with density-dependent and allogenic mortality both active at the same time. CWD volume was higher in plot Valbona 2a than in Valbona 1, but neither was comparable yet to the reference old-growth sites from Central Europe, both in quantity and in quality (e.g., decay rate continuity). The effects of the past management were: (1) reducing the quality and quantity of the CWD, (2) alleviating competition, (3) increasing resistance to minor disturbances and, as a consequence, (4) delaying the development processes. In mature or overmature subalpine Norway spruce stands withdrawn from regular management many decades are necessary to develop old-growth characteristics and a longer period of time is necessary to reach a true old-growth stage

Development of old-growth characteristics in uneven-aged forests of the Italian Alps

During the last millennia, all forests of the Italian Alps have been heavily affected by human land-use. Consequently, forest structures have been modified, and there are no old growth remains. In the last decades, however, many forests have been withdrawn from regular management, because wood production was unprofitable, and left to develop naturally. At the same time, in currently managed forests, silvicultural systems able to develop or maintain old-growth characteristics are being required. The aim of this paper was to assess the status and developmental dynamics of old-growth characteristics in mixed beech, silver fir, and Norway spruce montane forests of the eastern Italian Alps. We selected along a naturalness gradient (a) three old-growth forests in Bosnia and Montenegro (due to the lack of old-growth forests in the Italian Alps), (b) two forests withdrawn from regular management for at least 50 years, and (c) three currently managed forests. In each forest, we analysed 17 structural attributes, in order to assess their value as indicators of old-growth condition. Old-growth forests were characterized by significantly higher amounts of live and dead biomass, share of beech in the dominant and regeneration layers, and number of large trees. The diameter distribution was best described as a rotated sigmoid, differently from currently and formerly managed forest. We discuss the differences in old-growth characteristics across the management gradient and use our results to evaluate the effectiveness of retention prescriptions currently applied in the studied regions in maintaining or promoting old-growth structural attributes in managed forests

Scaling law in the Standard Map critical function. Interpolating hamiltonian and frequency map analysis

We study the behaviour of the Standard map critical function in a neighbourhood of a fixed resonance, that is the scaling law at the fixed resonance. We prove that for the fundamental resonance the scaling law is linear. We show numerical evidence that for the other resonances $p/q$, $q \geq 2$, $p \neq 0$ and $p$ and $q$ relatively prime, the scaling law follows a power--law with exponent $1/q$.Comment: AMS-LaTeX2e, 29 pages with 8 figures, submitted to Nonlinearit

Corso di Selvicoltura in Basilicata:un utile confronto fra gruppi di esperti.

Training course in silviculture: a useful comparison among expert group people. The results are presented of a training course in silviculture performed in experimental areas located in oak and beech woods in Basilicata (Southern Italy). In particular, the different choices made, in terms of thinning, by different groups of experts who worked independently in these areas are quantitatively summarized and discussed

Effects of forest management on ground beetle diversity in alpine beech (Fagus sylvatica L.) stands

European beech forests are of particular importance for biodiversity, although relatively little is known about how beech forest management impacts on invertebrate communities. In this paper we investigated the influence of beech forest management history [i.e. over-mature coppices (OC) and coppices in conversion to high forests (CCHF)], climatic, topographic and microhabitat characteristics on ground beetle diversity (measured as total relative abundance, species richness, Shannon diversity and abundance of the endangered endemic species Carabus olympiae) in northern Italy. The diversity of forest specialist carabids was higher in OC and in forest stands characterized by a higher mean temperature and lower relative humidity. Moreover, we detected a positive response of several diversity variables to coarse wood debris cover or volume, herb cover, and the standard deviation of tree diameter. Currently, OC seems to be a more favorable habitat for forest carabids, including C. olympiae, although succession over time can lead to a progressive homogenization of the vegetation structure, with negative consequences for the conservation of the forest carabid assemblage.Based on our results, we suggest that the traditional management of beech coppice and its conversion to high forest be modified by including practices aimed at promoting structural and microhabitat diversity such as retention of large trees, creation of canopy gaps, retention of coarse wood debris and the preservation of 'islands' of older trees in the managed stands

Voluntary carbon credits from improved forest management : policy guidelines and case study

Human activities have the potential to enhance carbon sequestration by the world\ue2\u80\u99s forests and contribute to climate change mitigation. Voluntary carbon trading is currently the only option to pursue and reward carbon sequestration by forestry activities. Carbon credits for enhanced sequestration can be sold to partners wishing to offset their own emissions. Here we illustrate the steps taken to design guidelines for the generation of voluntary carbon credits by improved forest management in Piemonte, Italy. The guidelines have been developed in a joint effort by academia, regional administrations, forest owners and professional consultants. In particular, we show how to compute the baseline and the additionality of credit-generating forest management activities, and how to reconcile the generation of forest carbon credits with law requirements, technical limitations, and the provision of other ecosystem services. To illustrate the profitability of carbon credit generation, we simulated the application of carbon credit guidelines to two forest-rich mountain watersheds in the southern part of the Piemonte region. The two dominating tree species are beech (Fagus sylvatica L.) and chestnut (Castanea sativa Mill.). We computed current forest carbon stock and carbon credits generated in 20 years under business as usual and an alternative biomass retention scenario. The IFM resulted in an avoided harvest of 39,362 m3for a net total of 64,014 MgCO2e after subtracting harvest emissions, or 38 Mg ha-1throughout the permanence period of 20 years. These steps can be replicated in other mountain regions where there is interest in promoting this ecosystem service as an alternative or an addition to production-oriented forest management

Drivers of Pinus sylvestris L. regeneration following small, high-severity fire in a dry, inner-alpine valley

In dry, inner-alpine valleys, regeneration of Scots pine after large, high-severity fires is limited by seed dispersal distance. When dispersal limitations are relaxed (small fires), colonization dynamics of Scots pine remain unclear. Our aims were (1) to assess the regeneration niche of Scots pine seedlings following small fires in a dry, inner-alpine valley and (2) to measure the influence of competition, drought, fertility, and soil pH on pine colonization. We sampled pine seedlings in six high-severity fires (1962\ue2\u80\u932006), where distance from the unburned edge was <\uc2\ua060\uc2\ua0m. We modeled seedling density as a function of topography and soil cover by means of generalized linear models. Ellenberg indicator values (EIVs) for light, moisture, soil pH, and fertility were computed from vegetation sampling at each plot, and used as additional predictors of seedling density. Pine colonization was initially slow due to drought, and peaked 16 years after fire. The inclusion of EIV provided +31% of explained deviance in models of seedling density. Herb cover and soil moisture were the most important predictors. Scots pine regenerates successfully following small, high-severity fires, albeit slowly due to unfavorable water balance. Hence, restoration by artificial regeneration may not be necessary when seed dispersal is not limiting