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Possible origins of macroscopic left-right asymmetry in organisms
I consider the microscopic mechanisms by which a particular left-right (L/R)
asymmetry is generated at the organism level from the microscopic handedness of
cytoskeletal molecules. In light of a fundamental symmetry principle, the
typical pattern-formation mechanisms of diffusion plus regulation cannot
implement the "right-hand rule"; at the microscopic level, the cell's
cytoskeleton of chiral filaments seems always to be involved, usually in
collective states driven by polymerization forces or molecular motors. It seems
particularly easy for handedness to emerge in a shear or rotation in the
background of an effectively two-dimensional system, such as the cell membrane
or a layer of cells, as this requires no pre-existing axis apart from the layer
normal. I detail a scenario involving actin/myosin layers in snails and in C.
elegans, and also one about the microtubule layer in plant cells. I also survey
the other examples that I am aware of, such as the emergence of handedness such
as the emergence of handedness in neurons, in eukaryote cell motility, and in
non-flagellated bacteria.Comment: 42 pages, 6 figures, resubmitted to J. Stat. Phys. special issue.
Major rewrite, rearranged sections/subsections, new Fig 3 + 6, new physics in
Sec 2.4 and 3.4.1, added Sec 5 and subsections of Sec
Transverse Spin Asymmetries in Drell-Yan Processes: }
We consider two different spin asymmetries in Drell-Yan processes generated
by the collisions of an unpolarized proton beam on transversely polarized
protons: the muon helicity asymmetry and the left-right asymmetry. We calculate
the asymmetries in the framework of the QCD improved parton model, taking into
account the parton transverse momentum and considering the first order QCD
corrections. The muon helicity asymmetry is sensitive to the quark transversity
distribution and is nonvanishing even at zero order. On the contrary the
left-right asymmetry vanishes at zero order but not at first order in the QCD
coupling constant, as a result of gluon contribution
Lambda polarization and single-spin left-right asymmetry in diffractive hadron-hadron collisions
We discuss Lambda polarization and single-spin left-right asymmetry in
diffractive hadron-hadron scattering at high energies. We show that the
physical picture proposed in a recent Letter is consistent with the
experimental observation that polarization in the diffractive
process, , is much higher than that in the inclusive
reaction, . We make predictions for the left-right asymmetry,
A_N, and for the spin transfer, , in the single-spin process
and suggest further experimental tests of the
proposed picture.Comment: 14 pages, 3 ps-figures. Phys. Rev. D (in press
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