Using abundance data to assess the relative role of sampling biases and evolutionary radiations in Upper Muschelkalk ammonoids

The Middle Triassic ammonoid genus Ceratites diversified spectacularly within the Germanic Muschelkalk Basin during the Anisian/Ladian (244–232 Mya). Previous studies have interpreted this diversification as a sequence of rapid, endemic radiations from a few immigrant taxa. Here we investigate the possibility that geological and sampling biases, rather than ecological and evolutionary processes, are responsible for this pattern. A new specimen based dataset of Ceratites species-richness and abundance was assembled. This dataset was combined with 1:200000 geological maps in a geodatabase to facilitate geospatial analyses. One set of analyses compared species richness per geological map with the number of occurrences and localities per map. Per-map change in the amount of rock available to sample for fossils was also included as a variable. Of these three variables, number of occurrences is the most strongly correlated with richness. Variation in the amount of rock is not a strong determinant of species-richness. However, rarefaction of basin-wide species/abundance data demonstrates that differences in species-richness through time are not attributable to sample size differences. The average percent similarity among sites remained close to 50% throughout the Upper Muschelkalk. The rank abundance distribution (RAD) of species from the first interval of the Upper Muschelkalk is consistent with colonization of a disturbed environment, while the other two intervals have RADs consistent with more stable ecosystems. These results indicate that genuine ecological and evolutionary events are partly responsible for the observed differences in richness and abundance. Although changes in the RADs through time support changes in the ammonoid assemblage structure, the processes underlying increasing richness and change in RADS cannot be explained by increasing geographic distinctiveness or isolation among the ammonoid assemblages present at different localities

The Carnian Humid Episode of the late Triassic: a review

From 1989 to 1994 a series of papers outlined evidence for a brief episode of climate change from arid to humid, and then back to arid, during the Carnian Stage of the late Triassic Epoch. This time of climate change was compared to marine and terrestrial biotic changes, mainly extinction and then radiation of flora and fauna. Subsequently termed, albeit incorrectly, the Carnian Pluvial Event (CPE) by successive authors, interest in this episode of climatic change has increased steadily, with new evidence being published as well as several challenges to the theory. The exact nature of this humid episode, whether reflecting widespread precipitation or more local effects, as well as its ultimate cause, remains equivocal. Bed-by-bed sampling of the Carnian in the Southern Alps (Dolomites) shows the episode began with a negative carbon isotope excursion that lasted for only part of one ammonoid zone (A. austriacum). However, that the Carnian Humid Episode represents a significantly longer period, both environmentally and biotically, is irrefutable. The evidence is strongest in the European, Middle Eastern, Himalayan, North American and Japanese successions, but not always so clear in South America, Antarctica and Australia. The eruption of the Wrangellia Large Igneous Province and global warming (causing increased evaporation in the Tethyan and Panthalassic oceans) are suggested as causes for the humid episode

The Permian-Triassic boundary and Early Triassic sedimentation in Western European basins: an overview

At the scale of the peri-Tethyan basins of western Europe, the “Buntsandstein” continental lithostratigraphic units are frequently attributed to the “Permian-Triassic” because, in most cases, the lack of any “Scythian” (i.e. Early Triassic) biochronological evidence makes it very difficult to attribute the basal beds of the cycle to the Permian or to the Triassic. A careful recognition of unconformities and sedimentary indications of clearly arid climate provide powerful tools for correlation within non-marine successions that are devoid of any biostratigraphic markers, at least on the scale of the West European Plate. From a review of the “Buntsandstein” series of several basins we can characterize the Permian-Triassic boundary and the beginning of Triassic sedimentation at the scale of Western Europe. We clearly show that, except for the Central Germanic Basin, an unconformity can be observed between the Permian and the Triassic. Apart from the Germanic Basin, there is a total lack of typically “Scythian” fossils in the rest of West European basins, and the oldest biochronological markers yielded by these units are palynomorphs allowing to assign an Anisian age generally to the upper part of the “Buntsandstein”, but also its lowermost in a few cases. In the peri-Tethyan basins of western Europe, the Permian-Triassic boundary corresponds to an unconformity overlain by conglomerates containing ventifacts (followed by fluvial sandstones, sometimes rich in paleosols and sometimes totally devoid), which are attributed mainly to the lower Olenekian, i.e. Smithian. Alternatively, the succession passes up directly into fluvial sandstones containing the first paleosols, and then plant debris and palynomorphs attributed to the Anisian. In this way, the lack of typically Early Triassic fossils in most of the peri-Tethyan basins, at the scale of the west European Plate, can be explained by a true stratigraphic hiatus in the earliest Triassic (i.e. Induan) and by arid conditions unfavourable for the development of flora and fauna and their preservation during the Olenekian.En las cuencas peritéthicas de Europa occidental, la unidad litoestratigráfi ca continental “Buntsandstein” es frecuentemente atribuida al “Pérmico-Triásico”, porque, en la mayoría de los casos, la falta de elementos biocronológicos “scythienses” complica la atribución de los niveles basales al Pérmico o al Triásico. Un cuidadoso examen de las discordancias, y el uso de indicadores sedimentarios de climas claramente áridos constituyen herramientas fundamentales para la correlación de estas unidades no marinas, desprovistas de cualquier marcador bioestratigráfi co, al menos en el Oeste de la Placa Europea. La revisión de las series “Buntsandstein” de varias cuencas nos ha permitido caracterizar el límite Pérmico-Triásico y el comienzo de la sedimentación durante el Triásico inferior en el oeste de Europa, demostrando claramente que, excepto en la Cuenca Germánica Central, se puede observar una discordancia entre las series pérmicas y triásicas. Excepto en esta Cuenca Germánica, existe una falta total de fósiles típicamente “scythienses”, y los elementos bioestratigráfi cos más antiguos encontrados son conjuntos de palinomorfos que permiten asignar a los niveles superiores de las series “Buntsandstein” una edad Anisiense, en algunos cortes los niveles inferiores. En las cuencas peritéthicas de Europa occidental, el límite Pérmico-Triásico se corresponde con una discordancia que se recubre por conglomerados con ventifactos (seguidos por areniscas fl uviales, a veces ricas en paleosuelos y a veces totalmente desprovistas de ellos), atribuidos principalmente al Olenekiense inferior (Smithiense), o directamente areniscas fluviales donde aparecen los primeros paleosuelos, así como restos de plantas y palinomorfos, atribuidos al Anisiense. De esta forma en el Oeste de la Placa Europea, la falta de fósiles característicos del Triásico Inferior, en la mayoría de las cuencas perithéticas, puede explicarse por un importante hiato estratigráfi co durante el Triásico Inferior (Induense) y por unas condiciones áridas, desfavorables para la vida y la preservación de fósiles durante el Olenekiense

International field workshop on "The Triassic of Eastern France", october 2-7, 2006

Mémoire de Géosciences-Rennes, Hors-série n°5Excursion géologique organisée à l'occasion du congrès international "The triasic of Eastern France", du 2 au 7 ocrobre 200

High-resolution early Mesozoic Pangean climatic transect in lacustrine environments

Analysis of 6700 m of core from the Newark rift basin in New Jersey, USA provides a high-resolution astronomically calibrated magnetic polarity time scale for the Late Triassic and Early Jurassic spanning about 33 million years. This time scale, and its application elsewhere, allows a significant simplification of the pattern of climate-sensitive facies in the early Mesozoic basins of the central and north Atlantic margins. Coals and deep-water lacustrine deposits were produced at the paleoequator (Richmond-type sequences), while strikingly cyclical lacustrine and playa deposits were produced \00 to the north and south (Newark-type lacustrine sequences). At 10-30 ON, eolian dunes, playas sediments and evaporites were deposited (Fundy-type sequences). Farther north, shallow-water lacustrine red beds were deposited (Fleming Fjord-type sequences), while yet farther north (-40°), perennial-lake black mudstones and coals again dominated in the humid temperate zone (Kap Stewart-type sequences). Central Pangea drifted north about 10° during the Late Triassic, and the vertical sequence of climate-sensitive facies in individual basins changed as the basins passed through different climate zones. This simple zonal climate pattern explains most first-order changes in overall lacustrine sequences seen in the rift zone. Lakelevel cycles of Milankovitch origin change in a predicable way with the latitudinal shifts in climate and lacustrine style. Roughly \0 ky precessional cycles dominate within a few degrees of the equator, while -20 ky precessional cycles are dominant northward to about 30 ON where 40 ky obliquity cycles become evident in lake-level records

Co-occurrence of Neusticosaurus edwardsii and N. peyeri (Reptilia) in the Lower Meride Limestone (Middle Triassic, Monte San Giorgio)

A newly opened excavation in the Cassina beds of the Lower Meride Limestone (Monte San Giorgio UNESCO World Heritage List, Canton Ticino, Switzerland), has yielded a pachypleurosaurid (Reptilia: Sauropterygia) specimen which is identified as Neusticosaurus peyeri. The resulting co-occurrence of N. peyeri and N. edwardsii, the latter so far regarded as the sole species of the genus present in this horizon, challenges the hypothesis of a single anagenetic lineage in Neusticosaurus species from Monte San Giorgio. In addition, it leads to a reconsideration of the phylogenetic inferences about Neusticosaurus evolution in the Monte San Giorgio area. The stratigraphic distribution of the Neusticosaurus species in the Monte San Giorgio basin is updated on the basis of recent find

Climate warming, euxinia and carbon isotope perturbations during the Carnian (Triassic) Crisis in South China

The Carnian Humid Episode (CHE), also known as the Carnian Pluvial Event, and associated biotic changes are major enigmas of the Mesozoic record in western Tethys. We show that the CHE also occurred in eastern Tethys (South China), suggestive of a much more widespread and probably global climate perturbation. Oxygen isotope records from conodont apatite indicate a double-pulse warming event. The CHE coincided with an initial warming of 4 °C. This was followed by a transient cooling period and then a prolonged ~7 °C warming in the later Carnian (Tuvalian 2). Carbon isotope perturbations associated with the CHE of western Tethys occurred contemporaneously in South China, and mark the start of a prolonged period of carbon cycle instability that persisted until the late Carnian. The dry-wet transition during the CHE coincides with the negative carbon isotope excursion and the temperature rise, pointing to an intensification of hydrologic cycle activities due to climatic warming. While carbonate platform shutdown in western Tethys is associated with an influx of siliciclastic sediment, the eastern Tethyan carbonate platforms are overlain by deep-water anoxic facies. The transition from oxygenated to euxinic facies was via a condensed, manganiferous carbonate (MnO content up to 15.1 wt%), that records an intense Mn shuttle operating in the basin. Significant siliciclastic influx in South China only occurred after the CHE climatic changes and was probably due to foreland basin development at the onset of the Indosinian Orogeny. The mid-Carnian biotic crisis thus coincided with several phenomena associated with major extinction events: a carbonate production crisis, climate warming, δ 13 C oscillations, marine anoxia, biotic turnover and flood basalt eruptions (of the Wrangellia Large Igneous Province)

Silicification and organic matter preservation in the Anisian Muschelkalk: Implications for the basin dynamics of the central European Muschelkalk Sea

Anisian Muschelkalk carbonates of the southern Germanic Basin containing silicified ooidal grainstone are interpreted as evidence of changing pH conditions triggered by increased bioproductivity (marine phytoplankton) and terrestrial input of plant debris during maximum flooding. Three distinct stages of calcite ooid replacement by silica were detected. Stage 1 reflects authigenic quartz development during the growth of the ooids, suggesting a change in the pH–temperature regime of the depositional environment. Stages 2 and 3 are found in silica-rich domains. The composition of silica-rich ooids shows significant Al2O3 and SrO but no FeO and MnO, indicating that late diagenetic alteration was minor. Silicified interparticle pore space is characterized by excellent preservation of marine prasinophytes; palynological slides show high abundance of terrestrial phytoclasts. The implications of our findings for basin dynamics reach from paleogeography to cyclostratigraphy and sequence stratigraphy, since changes in the seawater chemistry and sedimentary organic matter distribution reflect both the marine conditions as well as the hinterland. Basin interior changes might overprint the influence of the Tethys Ocean through the eastern and western gate areas. Stratigraphically, such changes might enhance marine flooding signals. Ongoing research needs to address the complex interaction between an intracratonic basin and an open-ocean system by comparing local and regional biotic and abiotic signals